| Citation | Magtoon, W. and R. Arai, 1993, Karyotypes and distribution of nucleolus organizer regions in cyprinid fishes from Thailand.. Jap. J. Ichthyol. 40(1):77-85. |
| Ref. | Magtoon, W. and R. Arai, 1993 |
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| Named Used as Valid | Puntioplites proctozysron |
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