| 引用出处 | Berg, L.S., 1965, Freshwater fishes of the U.S.S.R. and adjacent countries. volume 3, 4th edition.. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1949). |
| 参考文献 | Berg, L.S., 1965 |
| 页 | 38-42 |
| 命名为有效 | Gasterosteus aculeatus |
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