| 引用出处 | Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov, 1997, An annotated check-list of the freshwater fishes of Russia.. J. Ichthyol. 37(9):687-736. |
| 参考文献 | Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov, 1997 |
| 页 | 711 |
| 命名为有效 | Gasterosteus aculeatus |
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