| 引用出處 | Taki, Y., T. Urushido, A. Suzuki and C. Serizawa, 1977, A comparative chromosome study of Puntius (Cyprinidae: Pisces). I. Southeast Asian species.. Proc. Japan Acad., Ser. B: Phys. Biol. Sci. 53(6):231-235. |
| 參考文獻 | Taki, Y., T. Urushido, A. Suzuki and C. Serizawa, 1977 |
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| 已命名為有效 | Puntius titteya |
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