| 引用出处 | Economidis, P.S., E. Dimitriou, R. Pagoni, E. Michaloudi and L. Natsis, 2000, Introduced and translocated fish species in the inland waters of Greece.. Fish. Manage. Ecol. 7:239-250. |
| 参考文献 | Economidis, P.S., E. Dimitriou, R. Pagoni, E. Michaloudi and L. Natsis, 2000 |
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| 命名为有效 | Gasterosteus aculeatus |
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