| 引用出处 | Pietsch, T.W., K. Amaoka, D.E. Stevenson, E.L. MacDonald, B.K. Urbain and J.A. López, 2001, Freshwater fishes of the Kuril Islands and adjacent regions.. Species Diversity 6:133-164. |
| 参考文献 | Pietsch, T.W., K. Amaoka, D.E. Stevenson, E.L. MacDonald, B.K. Urbain and J.A. López, 2001 |
| 页 | 9 |
| 命名为有效 | Gasterosteus aculeatus aculeatus |
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