| 引用出处 | Arnoult, J., 1986, Gasterosteidae.. p. 280. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. |
| 参考文献 | Arnoult, J., 1986 |
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| 命名为有效 | Gasterosteus aculeatus |
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