| 引用出处 | Banister, K., 1986, Gasterosteidae.. p. 640-643. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the north-eastern Atlantic and the Mediterranean. volume 2. UNESCO, Paris. |
| 参考文献 | Banister, K., 1986 |
| 页 | 640 |
| 命名为有效 | Gasterosteus aculeatus |
| 附注 | Fig., map. |
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