| Kutipan | Ishihara, H., 1990, Study on the systematics and fishery resources of the North Pacific skates (Pisces: Chondrichthyes: Rajidae).. Tokyo University. 186 p. Ph.D. dissertation. |
| Acuan | Ishihara, H., 1990 |
| Halaman | |
| Bernama Digunakan sebagai Valid | Pavoraja asperula |
| Bagaimana cara | Cited in Ref. 41250. Not actually seen. Name used as valid appeared in this reference. |
| Lokalitas | |
| Penyebaran | |
| Kutipan |