| 引用出处 | Whoriskey, F.G. and R.J. Wootton, 1987, The swimming endurance of threespine sticklebacks, Gasterosteus aculeatus L., from the Afon Rheidol, Wales.. J. Fish Biol. 30(3):335-339. |
| 参考文献 | Whoriskey, F.G. and R.J. Wootton, 1987 |
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| 命名为有效 | Gasterosteus aculeatus |
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