| 引用出处 | Witkowski, A., J. Blachuta, B. Kokurewicz and M. Kowalewski, 1989, Changes of gonadosomatic intex (GSI) and egg diameter in the grayling Thymallus thymallus (L.) in annual cycle.. Acta Ichthyol. Piscat. 19(2):21-28. |
| 参考文献 | Witkowski, A., J. Blachuta, B. Kokurewicz and M. Kowalewski, 1989 |
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| 命名为有效 | Thymallus thymallus |
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