| 引用出处 | Elvira, B., 1996, Endangered freshwater fish of Spain.. p. 55-61. A. Kirchhofer and D. Hefti (eds) In Conservation of Endangered Freshwater Fish in Europe. Birkhäuser Verlag, Basel, Switzerland. |
| 参考文献 | Elvira, B., 1996 |
| 页 | 57 |
| 命名为有效 | Gasterosteus aculeatus |
| 附注 | |
| 地点 | |
| 分布 | Spain |
| 报价 |