| 引用出处 | Mikheev, P.B., V.I. Ostrovskii, N.N. Semenchenko, G.V. Novomodnyi, A.P. Shmigirilov, A.L. Antonov and E.I. Barabanshchikov, 2012, Biological Specific Features of the Lower Amur Grayling Thymallus tugarinae (Salmoniformes: Thymallidae): 1. Age Composition.. J. Ichthyol. 52(8):519-527. |
| 参考文献 | Mikheev, P.B., V.I. Ostrovskii, N.N. Semenchenko, G.V. Novomodnyi, A.P. Shmigirilov, A.L. Antonov and E.I. Barabanshchikov, 2012 |
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| 命名为有效 | Thymallus tugarinae |
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