| 引用出处 | Akita Prefecture, 2002, Threatened Wildlife of Akita Prefecture, Animals.. Akita Prefecture, Akita. (http://www.pref.akita.lg.jp/www/contents/1135148036456/; http://www.pref.akita.lg.jp/www/contents/1135148036456/files/fishes.pdf) |
| 参考文献 | Akita Prefecture, 2002 |
| 页 | |
| 命名为有效 | Gasterosteus aculeatus aculeatus |
| 附注 | |
| 地点 | |
| 分布 | Japan (Tohoku: Akita) |
| 报价 |