| 引用出处 | Niigata Prefecture, 2001, Red Data Book of Niigata.. Niigata Prefecture, Niigata. (http://www.pref.niigata.lg.jp/kankyokikaku/1214240790991.html; http://www.pref.niigata.lg.jp/HTML_Article/rdb12_gyorui.pdf) |
| 参考文献 | Niigata Prefecture, 2001 |
| 页 | |
| 命名为有效 | Gasterosteus aculeatus aculeatus |
| 附注 | |
| 地点 | |
| 分布 | Japan (Chubu: Niigata) |
| 报价 |