Upper part of the base of the caudal fin with one elongated dark spot; broad axe-shaped lamina Canestrini in males; scales more or les rounded with relatively small central part devoid of cross furrows; Gambetta's zones of pigmentation well developed, the fourth zone consisting of relative large 13-17, usually 16 spots with depth equals or exceeds (the largest spots before dorsal fin) horizontal eye diameter; suborbital spine in females not reaching posterior edge of pupil, but in smaller males reaches as far as the posterior edge of the pupil or some more; the outer branch of this spine usually (85.7%) not reaching the center of the eye; head some shortened and deep with more or less straight profile; karyotype consists of 50 (10 meta-, 30 submeta-, 8 subtelo- and 2 areocentric) chromosomes, the number of chromosomes arms (NF) - 90 (Ref. 57856).