This fine-scaled species have the following characteristics: pyloric caeca usually 5-6, typically 5 + 1; usually 3 rudimentary gill rakers on the upper arm of the first arch and 4-5 developed rakers on lower arm with 2-5 rudimentary rakers anteriorly, total gill rakers usually 11-12 (10-15); count of scale rows almost 30 between the occipital line and the dorsal-fin origin, 14-15 rows between the dorsal-fin origin and the lateral line, and about 255 lateral scale rows; dorsal-fin origin between neural spines 4-5 (rarely between 3-4); color of head and body rather uniformly dusky, silvery, or whitish along the belly; dorsal fin usually lacks a dark edge, and when present it is poorly developed; anal fin has an inconspicuous and narrow dark edge; orobranchial cavity usually pale, but the rear of the roof of the mouth may be dusky; esophagus blackish; stomach and hindgut pale (Ref. 91765).