Females: esca with a low, distal prolongation of bulb, length about half diameter of bulb, distally subdivided into two blunt tips (posterior tip the larger); on each side of base of distal prolongation a single, compressed filament, length about three times diameter of bulb, with 2-4 short, pointed, side-branches; a low papilliform posterior appendage behind escal pore. Barbel about 25% SL in total length, distally divided at about half length into three main-branches, and, at about half distance between this division and base, with a posterior pair of shorter side-branches; all branches repeatedly divided, each terminal branch with a series of internal photophores and one or more distal filaments. Dense concentration of subdermal pigment on caudal peduncle (Ref. 42949).