| Main Ref. | Kottelat, M. and H.H. Tan, 2008 |
| Appearance refers to | |
| Bones in OsteoBase |
| Specialized organs | |
| Different appearance | |
| Different colors | |
| Remarks |
| Striking features | |
| Body shape lateral (shape guide) | |
| Cross section | |
| Dorsal head profile | |
| Type of eyes | |
| Type of mouth/snout | |
| Position of mouth | sub-terminal/inferior |
| Type of scales | |
| Diagnosis |
Diagnosed from all congeners from Java, Sumatra and Borneo by having 4½ scale rows between lateral line and dorsal-fin origin and 12 scales around the caudal peduncle. Can be further separated by having the combination of the following features: lateral line scales 32+2; 4 barbels; pectoral fin extending to 3/4 of distance to pelvic -fin origin; pelvic fin extending anal-fin origin; dorsal-fin tip extending slightly beyond vertical through anal-fin origin, when folded backwards; length of head less than body depth at dorsal-fin origin (Ref. 75626). Description: D iii,8½, the last two rays articulating on same pterygiophore ; A iii,5½ , the last two rays articulating on same pterygiophore; P i+15; V i+8 (Ref. 75626). |
| Ease of Identification |
| Lateral Lines | Interrupted: No |
| Scales on lateral line | 32 |
| Pored lateral line scales | |
| Scales in lateral series | |
| Scale rows above lateral line | 4 |
| Scale rows below lateral line | 4 |
| Scales around caudal peduncle | 12 |
| Barbels | 4 |
| Gill clefts (sharks/rays only) | |
| Gill rakers | |
| on lower limb | |
| on upper limb | |
| total | |
| Vertebrae | |
| preanal | |
| total |
| Attributes | no striking attributes |
| Fins number | 1 |
| Finlets No. | Dorsal |
| Ventral | |
| Spines total | |
| Soft-rays total | 11 - 11 |
| Adipose fin | absent |
| Attributes | forked |
| Fins number | |
| Spines total | |
| Soft-rays total | 8 - 8 |
| Pectoral | Attributes more or less normal |
| Spines | |
| Soft-rays 16 - 16 | |
| Pelvics | Attributes more or less normal |
| Position abdominal behind origin of D1 | |
| Spines | |
| Soft-rays 9 - 9 |