This species is distinguished by the following characters: two halves of the air bladder capsules connected by a manubrium, with posterior chamber free, well-developed, not encapsulated and more or less oval. In males, the first branched pectoral-fin ray is falcate, about twice the thickness of the second, branched once only near the tip and without a membrane between the branches except near the tip; other branched rays are also branched only once; branched ray 3-4 are the longest; a thick unculiferous pad on the dorsal surface of pectoral-fin rays, all rays and pads are covered dorsally by small, conspicuous, densely-set conical tubercles; suborbital flap spoon-shaped, with small tubercles on its posterior edge; no such modifications of pectoral fin in females (Ref. 132199)