This species is distinguished by the following characters: horizontal process of anguloarticular broadly covered by dentary, reaching posterior border of Meckelian cartilage (161:0 > 1); maxillary teeth extending across almost entire maxillary lamella (191:0 > 1); no hemal prezygapophysis on anterior caudal vertebrae (333:0 > 1); ventral margin of post-temporal anterior to lateral margin of the epioccipital (335:1 > 0; also in data set B as 266:1 > 0); lateral lamella of pelvic bone reaching two thirds of this bone (373:0 > 1); mandibular accessory tendon longitudinally inserted on vertical through middle or anterior half of Meckelian cartilage (474:0 > 1); gill filaments not fused to form a gill gland in adult males (511:1 > 0); and from the data set B, anterior margin of nasal extending anterior to lateral process of mesethmoid (14:1 > 0); parietal branch of supratemporal canal extend over frontal or not reaching lateral surface of parietal bone (33:0 > 1); first and second postcleithra separated from each other by distance greater than three-quarters of third postcleithrum length (296:1 > 0); dorsal margin of third postcleithrum not reaching second-postcleithrum midpoint (298:1 > 0); ventralmost tendinous fiber of hypaxialis inserted on the lateral or ventral surface of caudal-fin ray 19 (367:1 > 0); no terminal lateral-line tube on caudal-fin membrane (374:1 > 0); no humeral mark (388:1 > 0); distance between bases of middle caudal-fin rays 10 and 11 approximately equal or less than that between rays 11 and 12 (433:1 > 0); anal-fin bony hooks not confined to anterior half of fin (460:1 > 0) but, instead, distributing over proximalmost segment of all or, most, fin rays (462:0 > 1; 465:0 > 1) (Ref. 121996).