Classification List

Fish species numbers in the FishBase classification list


     The classification schema above genus level is limited in FishBase to classes, orders, families and subfamilies ranks (the latter not yet displayed). It has the advantage to keep it simple; it has the disadvantage to ignore some important clades in fish phylogeny like Chondrichthyes, Teleostei, Acanthomorpha, Percomorpha.

     In general, it follows Nelson's Fishes of the World (2006) with the noticeable exception of Elasmobranchii and Holocephali being elevated to class rank But at family level, FishBase may follow Eschmeyer's Catalog of Fishes that is updated more frequently to integrate new stable groupings.

     Number of taxa from Orders to Species are given for what they are, a useful tool for information and database management. The design of some analyses (so-called Biodiversity analyses) that uses these numbers must be carefully assessed before processing (see footnote for some critical points).

     Catalog of Fishes gives also number of species, including available names in the sense of the International Code of Zoological Nomenclature, which we don't have. Possible discrepancies between the species and genera numbers are explained in Bailly (2010). As Catalog of Fishes is updated ahead of FishBase, the numbers given there are in general more accurate.

     Wiley and Johnson published a new classification for Teleostei (2010). They propose to stabilize several hypotheses published recently or some (long) time ago by listing the known evidences (synapomorphies). FishBase will follow this new arrangement as soon as the ichthyological community endorses it, e.g., as published in a next version of Fishes of the World. In the meantime, to avoid confusion for non-specialists, we hold on the current one.

Note that the numbers of species given below does not take into account subspecies.

Total count of Genera: 5276
Total count of Species: 36761


Number of Valid
Kingdom Phylum
Class
#6
Order
#93
Family
#629
Orders Families Genera Species
Animalia
Chordata
Myxini 1 1 6 92
Myxiniformes 1 6 92
Myxinidae 6 92
Petromyzonti 1 3 11 50
Petromyzontiformes 3 11 50
Petromyzontidae 9 45
Geotriidae 1 2
Mordaciidae 1 3
Elasmobranchii 13 65 214 1299
Hexanchiformes 2 4 7
Hexanchidae 3 5
Chlamydoselachidae 1 2
Heterodontiformes 1 1 10
Heterodontidae 1 10
Orectolobiformes 7 13 45
Rhincodontidae 1 1
Parascylliidae 2 8
Brachaeluridae 1 2
Orectolobidae 3 12
Hemiscylliidae 2 17
Stegostomatidae 1 1
Ginglymostomatidae 3 4
Lamniformes 8 10 16
Mitsukurinidae 1 1
Carchariidae 1 2
Odontaspididae 1 2
Pseudocarchariidae 1 1
Megachasmidae 1 1
Alopiidae 1 3
Cetorhinidae 1 1
Lamnidae 3 5
Carcharhiniformes 12 53 309
Pentanchidae 11 111
Dichichthyidae 1 5
Scyliorhinidae 4 39
Atelomycteridae 3 13
Proscylliidae 3 7
Pseudotriakidae 3 5
Leptochariidae 1 1
Triakidae 9 47
Hemigaleidae 4 8
Carcharhinidae 11 59
Galeocerdonidae 1 1
Sphyrnidae 2 13
Squaliformes 6 22 144
Dalatiidae 7 10
Etmopteridae 4 54
Somniosidae 6 17
Oxynotidae 1 5
Centrophoridae 2 17
Squalidae 2 41
Echinorhiniformes 1 1 2
Echinorhinidae 1 2
Pristiophoriformes 1 2 10
Pristiophoridae 2 10
Squatiniformes 1 1 26
Squatinidae 1 26
Torpediniformes 5 16 73
Platyrhinidae 2 5
Narkidae 6 12
Narcinidae 5 31
Hypnidae 1 1
Torpedinidae 2 24
Rhinopristiformes 5 13 76
Trygonorrhinidae 3 8
Rhinobatidae 4 40
Rhinidae 3 11
Glaucostegidae 1 10
Pristidae 2 7
Rajiformes 4 40 314
Rajidae 18 165
Arhynchobatidae 14 114
Gurgesiellidae 3 19
Anacanthobatidae 5 16
Myliobatiformes 12 38 267
Zanobatidae 1 2
Hexatrygonidae 1 1
Dasyatidae 19 112
Potamotrygonidae 5 45
Urotrygonidae 2 20
Gymnuridae 1 16
Plesiobatidae 1 1
Urolophidae 3 28
Aetobatidae 1 5
Myliobatidae 2 20
Rhinopteridae 1 7
Mobulidae 1 10
Holocephali 1 3 6 58
Chimaeriformes 3 6 58
Callorhinchidae 1 3
Chimaeridae 2 46
Rhinochimaeridae 3 9
Cladistii 1 1 2 14
Polypteriformes 1 2 14
Polypteridae 2 14
Chondrostei 1 2 5 29
Acipenseriformes 2 5 29
Acipenseridae 3 27
Polyodontidae 2 2
Holostei 2 2 3 9
Lepisosteiformes 1 2 7
Lepisosteidae 2 7
Amiiformes 1 1 2
Amiidae 1 2
Teleostei 71 548 5025 35202
Hiodontiformes 1 1 2
Hiodontidae 1 2
Osteoglossiformes 6 31 266
Pantodontidae 1 1
Osteoglossidae 2 6
Arapaimidae 2 6
Notopteridae 4 10
Mormyridae 21 242
Gymnarchidae 1 1
Elopiformes 2 2 9
Elopidae 1 7
Megalopidae 1 2
Albuliformes 1 3 13
Albulidae 3 13
Notacanthiformes 2 6 30
Halosauridae 3 16
Notacanthidae 3 14
Anguilliformes 16 157 1043
Protanguillidae 1 1
Synaphobranchidae 12 51
Heterenchelyidae 2 8
Myrocongridae 1 6
Muraenidae 16 225
Chlopsidae 8 25
Colocongridae 1 11
Derichthyidae 2 3
Ophichthidae 63 362
Muraenesocidae 6 15
Nettastomatidae 6 47
Congridae 30 232
Moringuidae 2 18
Nemichthyidae 3 9
Serrivomeridae 2 10
Anguillidae 2 20
Saccopharyngiformes 5 5 28
Cyematidae 1 1
Monognathidae 1 15
Neocyematidae 1 1
Eurypharyngidae 1 1
Saccopharyngidae 1 10
Clupeiformes 10 85 446
Denticipitidae 1 1
Spratelloididae 2 8
Clupeidae 7 15
Dorosomatidae 30 115
Ehiravidae 11 31
Alosidae 4 32
Dussumieriidae 2 16
Engraulidae 18 188
Chirocentridae 1 2
Pristigasteridae 9 38
Alepocephaliformes 2 33 143
Alepocephalidae 20 103
Platytroctidae 13 40
Gonorynchiformes 4 7 41
Chanidae 1 1
Gonorynchidae 1 5
Kneriidae 4 34
Phractolaemidae 1 1
Cypriniformes 23 514 4860
Gyrinocheilidae 1 3
Catostomidae 15 85
Botiidae 8 62
Vaillantellidae 1 3
Cobitidae 20 238
Barbuccidae 1 3
Gastromyzontidae 20 153
Serpenticobitidae 1 3
Balitoridae 17 112
Ellopostomatidae 1 2
Nemacheilidae 45 817
Paedocyprididae 1 3
Psilorhynchidae 1 30
Cyprinidae 158 1798
Sundadanionidae 2 9
Danionidae 39 377
Leptobarbidae 1 5
Xenocyprididae 45 162
Tincidae 1 1
Acheilognathidae 7 82
Gobionidae 29 222
Tanichthyidae 1 6
Leuciscidae 99 684
Characiformes 28 291 2313
Citharinidae 3 8
Distichodontidae 16 110
Crenuchidae 12 108
Alestidae 19 120
Lepidarchidae 2 2
Hepsetidae 1 6
Tarumaniidae 1 1
Erythrinidae 3 20
Parodontidae 3 32
Cynodontidae 3 8
Serrasalmidae 17 105
Hemiodontidae 5 31
Anostomidae 16 166
Chilodidae 2 8
Curimatidae 8 111
Prochilodontidae 3 21
Lebiasinidae 6 75
Ctenoluciidae 2 7
Chalceidae 1 7
Triportheidae 5 23
Gasteropelecidae 3 9
Bryconidae 4 49
Iguanodectidae 3 32
Acestrorhynchidae 7 26
Spintherobolidae 2 5
Stevardiidae 44 342
Characidae 38 207
Acestrorhamphidae 62 674
Gymnotiformes 5 34 274
Gymnotidae 2 45
Apteronotidae 15 99
Sternopygidae 6 62
Hypopomidae 6 38
Rhamphichthyidae 5 30
Siluriformes 41 515 4175
Nematogenyidae 1 1
Trichomycteridae 43 363
Callichthyidae 12 235
Loricariidae 115 1042
Scoloplacidae 1 6
Astroblepidae 1 81
Diplomystidae 2 7
Cetopsidae 5 43
Chacidae 1 4
Plotosidae 10 43
Ritidae 1 7
Ailiidae 5 17
Horabagridae 2 8
Bagridae 20 245
Auchenoglanididae 3 30
Akysidae 5 58
Amblycipitidae 4 44
Sisoridae 25 313
Pangasiidae 4 30
Siluridae 12 108
Kryptoglanidae 1 1
Aspredinidae 13 48
Auchenipteridae 26 128
Doradidae 31 97
Heptapteridae 23 228
Phreatobiidae 1 3
Pimelodidae 32 116
Pseudopimelodidae 7 53
Clariidae 16 118
Heteropneustidae 1 6
Ariidae 39 161
Anchariidae 2 6
Austroglanididae 1 3
Cranoglanididae 1 5
Ictaluridae 7 52
Lacantuniidae 1 1
Amphiliidae 13 102
Malapteruridae 2 21
Mochokidae 9 232
Claroteidae 8 62
Schilbeidae 9 47
Lepidogalaxiiformes 1 1 1
Lepidogalaxiidae 1 1
Argentiniformes 4 23 94
Argentinidae 2 28
Microstomatidae 3 21
Bathylagidae 8 23
Opisthoproctidae 10 22
Galaxiiformes 1 7 66
Galaxiidae 7 66
Esociformes 2 4 14
Esocidae 1 7
Umbridae 3 7
Salmoniformes 1 11 245
Salmonidae 11 245
Osmeriformes 4 17 42
Osmeridae 6 15
Plecoglossidae 1 1
Salangidae 7 20
Retropinnidae 3 6
Stomiiformes 4 52 447
Gonostomatidae 8 32
Sternoptychidae 10 79
Phosichthyidae 7 24
Stomiidae 27 312
Ateleopodiformes 1 4 14
Ateleopodidae 4 14
Aulopiformes 17 46 296
Aulopidae 4 16
Synodontidae 4 85
Pseudotrichonotidae 1 4
Paraulopidae 1 14
Chlorophthalmidae 2 21
Ipnopidae 5 31
Bathysauroididae 1 1
Scopelarchidae 5 18
Notosudidae 3 17
Giganturidae 1 2
Bathysauridae 1 2
Paralepididae 11 69
Anotopteridae 1 3
Evermannellidae 3 8
Omosudidae 1 1
Alepisauridae 1 2
Sudidae 1 2
Myctophiformes 2 37 257
Neoscopelidae 3 7
Myctophidae 34 250
Lampriformes 6 11 28
Lampridae 1 6
Veliferidae 2 2
Lophotidae 2 5
Radiicephalidae 1 2
Trachipteridae 3 10
Regalecidae 2 3
Percopsiformes 3 7 12
Percopsidae 1 2
Aphredoderidae 1 3
Amblyopsidae 5 7
Zeiformes 6 16 33
Cyttidae 1 3
Oreosomatidae 4 10
Parazenidae 3 4
Zeniontidae 3 7
Grammicolepididae 3 3
Zeidae 2 6
Stylephoriformes 1 1 1
Stylephoridae 1 1
Gadiformes 17 84 635
Bregmacerotidae 1 17
Phycidae 2 11
Gaidropsaridae 3 19
Lotidae 3 5
Gadidae 11 22
Ranicipitidae 1 1
Merlucciidae 2 17
Euclichthyidae 1 3
Muraenolepididae 2 9
Melanonidae 1 2
Trachyrincidae 4 9
Moridae 19 109
Macruronidae 1 4
Lyconidae 1 2
Bathygadidae 2 26
Steindachneriidae 1 1
Macrouridae 29 378
Polymixiiformes 1 1 12
Polymixiidae 1 12
Beryciformes 8 31 127
Berycidae 2 10
Melamphaidae 5 73
Stephanoberycidae 4 4
Hispidoberycidae 1 1
Gibberichthyidae 1 2
Rondeletiidae 1 2
Barbourisiidae 1 1
Cetomimidae 16 34
Trachichthyiformes 5 20 73
Monocentridae 2 5
Trachichthyidae 8 53
Anomalopidae 6 9
Diretmidae 3 4
Anoplogastridae 1 2
Holocentriformes 1 8 90
Holocentridae 8 90
Ophidiiformes 4 121 573
Ophidiidae 49 286
Carapidae 9 38
Bythitidae 38 135
Dinematichthyidae 25 114
Batrachoidiformes 1 23 83
Batrachoididae 23 83
Scombriformes 16 76 280
Amarsipidae 1 1
Nomeidae 3 16
Ariommatidae 1 7
Stromateidae 3 17
Pomatomidae 1 1
Centrolophidae 7 31
Icosteidae 1 1
Arripidae 1 4
Tetragonuridae 1 3
Chiasmodontidae 4 32
Scombridae 15 55
Caristiidae 4 19
Bramidae 7 20
Scombrolabracidae 1 1
Gempylidae 16 26
Trichiuridae 10 46
Syngnathiformes 5 68 340
Aulostomidae 1 3
Fistulariidae 1 4
Centriscidae 5 12
Solenostomidae 1 7
Syngnathidae 60 314
Dactylopteriformes 2 4 14
Pegasidae 2 7
Dactylopteridae 2 7
Callionymiformes 2 18 213
Callionymidae 16 198
Draconettidae 2 15
Mulliformes 1 6 106
Mullidae 6 106
Kurtiformes 2 41 369
Kurtidae 1 2
Apogonidae 40 367
Gobiiformes 13 323 2422
Trichonotidae 1 10
Rhyacichthyidae 2 3
Odontobutidae 7 25
Oxudercidae 10 42
Eleotridae 22 148
Butidae 11 54
Milyeringidae 2 6
Thalasseleotrididae 3 4
Gobiidae 243 2001
Kraemeriidae 2 9
Microdesmidae 13 92
Xenisthmidae 6 16
Schindleriidae 1 12
Synbranchiformes 4 17 131
Mastacembelidae 3 90
Chaudhuriidae 6 10
Indostomidae 1 3
Synbranchidae 7 28
Anabantiformes 8 26 282
Anabantidae 4 33
Helostomatidae 1 1
Osphronemidae 14 138
Aenigmachannidae 1 1
Channidae 2 57
Nandidae 1 7
Badidae 2 37
Pristolepididae 1 8
Carangaria/misc 8 17 109
Latidae 3 14
Centropomidae 1 13
Lactariidae 1 1
Sphyraenidae 1 29
Leptobramidae 1 2
Toxotidae 1 7
Menidae 1 1
Polynemidae 8 42
Pleuronectiformes 16 130 823
Psettodidae 1 3
Citharidae 4 6
Scophthalmidae 4 9
Cyclopsettidae 4 46
Bothidae 20 173
Paralichthyidae 10 65
Pleuronectidae 25 62
Paralichthodidae 1 1
Oncopteridae 1 1
Rhombosoleidae 8 20
Achiropsettidae 4 4
Achiridae 7 35
Samaridae 4 30
Poecilopsettidae 3 21
Soleidae 31 184
Cynoglossidae 3 163
Carangiformes 7 51 173
Nematistiidae 1 1
Carangidae 39 147
Echeneidae 3 8
Rachycentridae 1 3
Coryphaenidae 1 2
Istiophoridae 5 11
Xiphiidae 1 1
Cichliformes 2 247 1805
Cichlidae 246 1803
Pholidichthyidae 1 2
Atheriniformes 11 51 384
Atherinopsidae 13 114
Atherinidae 13 69
Atherionidae 1 3
Bedotiidae 2 16
Isonidae 1 5
Melanotaeniidae 7 114
Phallostethidae 4 24
Dentatherinidae 1 1
Pseudomugilidae 3 19
Telmatherinidae 5 18
Notocheiridae 1 1
Cyprinodontiformes 14 153 1449
Aplocheilidae 2 14
Nothobranchiidae 12 309
Rivulidae 48 472
Pantanodontidae 4 10
Fundulidae 3 44
Cyprinodontidae 9 103
Profundulidae 2 13
Goodeidae 18 51
Fluviphylacidae 1 8
Poeciliidae 28 278
Anablepidae 3 20
Aphaniidae 8 45
Valenciidae 1 3
Procatopodidae 14 79
Beloniformes 6 34 299
Adrianichthyidae 2 45
Scomberesocidae 2 5
Belonidae 10 49
Exocoetidae 7 75
Hemiramphidae 8 61
Zenarchopteridae 5 64
Mugiliformes 1 26 78
Mugilidae 26 78
Gobiesociformes 1 53 189
Gobiesocidae 53 189
Blenniiformes 6 153 952
Tripterygiidae 29 185
Labrisomidae 16 123
Clinidae 26 92
Chaenopsidae 14 98
Dactyloscopidae 9 48
Blenniidae 59 406
Ovalentaria/misc 8 97 856
Ambassidae 9 56
Embiotocidae 13 24
Grammatidae 2 18
Opistognathidae 4 112
Plesiopidae 12 51
Polycentridae 4 5
Pomacentridae 29 434
Pseudochromidae 24 156
Acanthuriformes 13 54 453
Lobotidae 3 16
Pomacanthidae 7 91
Drepaneidae 1 3
Ephippidae 8 15
Chaetodontidae 12 136
Leiognathidae 10 53
Scatophagidae 2 4
Antigoniidae 1 18
Siganidae 1 29
Caproidae 1 1
Luvaridae 1 1
Zanclidae 1 1
Acanthuridae 6 85
Lophiiformes 21 74 405
Lophiidae 4 34
Antennariidae 6 32
Histiophrynidae 7 15
Tathicarpidae 1 1
Tetrabrachiidae 2 2
Lophichthyidae 1 1
Rhycheridae 1 2
Brachionichthyidae 5 14
Chaunacidae 2 33
Ogcocephalidae 10 93
Caulophrynidae 2 5
Neoceratiidae 1 1
Melanocetidae 1 6
Himantolophidae 1 22
Diceratiidae 2 7
Oneirodidae 17 77
Centrophrynidae 1 1
Ceratiidae 2 4
Thaumatichthyidae 1 3
Gigantactinidae 2 25
Linophrynidae 5 27
Tetraodontiformes 10 106 450
Triodontidae 1 1
Triacanthodidae 11 23
Triacanthidae 4 7
Aracanidae 6 14
Ostraciidae 6 25
Diodontidae 7 18
Tetraodontidae 29 206
Monacanthidae 27 109
Balistidae 12 42
Molidae 3 5
Centrarchiformes 20 78 301
Percichthyidae 9 24
Terapontidae 16 60
Kuhliidae 1 12
Dichistiidae 1 2
Oplegnathidae 1 7
Kyphosidae 2 16
Girellidae 2 21
Scorpididae 4 11
Microcanthidae 4 6
Parascorpididae 1 1
Centrarchidae 8 41
Elassomatidae 1 7
Enoplosidae 1 1
Sinipercidae 2 14
Aplodactylidae 1 5
Cheilodactylidae 1 2
Chironemidae 1 6
Cirrhitidae 12 33
Latridae 9 30
Percalatidae 1 2
Acropomatiformes 20 58 307
Scombropidae 1 3
Ostracoberycidae 1 3
Acropomatidae 2 13
Symphysanodontidae 2 13
Epigonidae 6 50
Howellidae 3 9
Polyprionidae 1 2
Lateolabracidae 1 2
Glaucosomatidae 1 4
Pempheridae 2 80
Dinolestidae 1 1
Stereolepididae 1 2
Banjosidae 1 3
Pentacerotidae 7 13
Malakichthyidae 3 17
Champsodontidae 1 13
Hemerocoetidae 10 31
Creediidae 8 20
Synagropidae 4 18
Bathyclupeidae 2 10
Eupercaria/misc 19 274 1807
Moronidae 2 6
Dinopercidae 2 2
Centrogenyidae 1 1
Callanthiidae 2 18
Priacanthidae 4 22
Sillaginidae 5 38
Malacanthidae 2 16
Latilidae 3 30
Emmelichthyidae 3 19
Lutjanidae 21 139
Gerreidae 8 53
Haemulidae 21 138
Sparidae 38 163
Lethrinidae 5 43
Nemipteridae 5 78
Sciaenidae 68 304
Monodactylidae 1 4
Cepolidae 3 47
Labridae 80 686
Perciformes/Percoidei 3 14 258
Percidae 11 248
Trachinidae 2 9
Niphonidae 1 1
Perciformes/Zoarcoidei 14 108 419
Zoarcidae 57 301
Anarhichadidae 2 5
Neozoarcidae 2 10
Eulophiidae 3 5
Stichaeidae 18 45
Lumpenidae 8 11
Opisthocentridae 6 12
Pholidae 4 15
Ptilichthyidae 1 1
Zaproridae 1 1
Cryptacanthodidae 1 4
Cebidichthyidae 1 1
Scytalinidae 1 1
Bathymasteridae 3 7
Perciformes/Bembropoidei 1 2 23
Bembropidae 2 23
Perciformes/Percophoidei 1 1 1
Percophidae 1 1
Perciformes/Notothenioidei 8 46 161
Bovichtidae 3 11
Pseudaphritidae 1 2
Eleginopidae 1 1
Nototheniidae 13 56
Harpagiferidae 1 12
Artedidraconidae 5 36
Bathydraconidae 11 18
Channichthyidae 11 25
Perciformes/Uranoscopoidei 6 27 198
Trichodontidae 2 2
Ammodytidae 7 34
Uranoscopidae 7 57
Leptoscopidae 3 5
Pinguipedidae 7 99
Cheimarrichthyidae 1 1
Perciformes/Serranoidei 6 81 604
Serranidae 12 106
Anthiadidae 38 250
Epinephelidae 16 172
Grammistidae 8 33
Liopropomatidae 6 42
Caesioscorpididae 1 1
Perciformes/Scorpaenoidei 21 136 847
Sebastidae 7 133
Setarchidae 4 11
Neosebastidae 2 18
Scorpaenidae 28 241
Apistidae 3 4
Tetrarogidae 17 44
Synanceiidae 9 40
Eschmeyeridae 1 1
Perryenidae 1 1
Aploactinidae 17 50
Pataecidae 3 3
Gnathanacanthidae 1 1
Congiopodidae 1 6
Zanclorhynchidae 2 3
Bembridae 5 12
Plectrogeniidae 1 2
Hoplichthyidae 1 14
Platycephalidae 18 84
Triglidae 8 132
Peristediidae 6 46
Normanichthyidae 1 1
Perciformes/Gasterosteoidei 3 8 24
Hypoptychidae 2 2
Gasterosteidae 5 21
Aulorhynchidae 1 1
Perciformes/Cottoidei 13 158 884
Anoplopomatidae 2 2
Zaniolepididae 2 3
Hexagrammidae 3 9
Rhamphocottidae 3 4
Jordaniidae 3 3
Cottidae 71 288
Psychrolutidae 10 40
Nautichthyidae 1 3
Hemilepidotidae 1 6
Hemitripteridae 2 5
Agonidae 21 46
Cyclopteridae 7 31
Liparidae 32 444
Coelacanthi 1 1 1 2
Coelacanthiformes 1 1 2
Latimeriidae 1 2
Dipneusti 1 3 3 6
Ceratodontiformes 3 3 6
Neoceratodontidae 1 1
Lepidosirenidae 1 1
Protopteridae 1 4


Footnote: Minelli makes a definitive point about ranking as being an arbitrary and subjective decision: "The main difficulty resides in determining whether two clades in distant parts of the phylogenetic tree deserve [being] acknowledged [at] the same rank or not." (Minelli, 2009:11). "But all this amounts to counting fruit by adding apples and cherries." (Minelli, 2009:12). In other words, if counting orders, families, and genera in a narrow context (e.g., number of genera in subfamilies of the same family) may be locally sensible (within a phylogenetic tree / classification), counting families across classes or orders is most often irrelevant: for instance, why assigning clown-fishes only the subfamily rank (Pomacentridae: Amphiprioninae) while the Moorish idol has a family of its own (Zanclidae), and not a subfamily in Acanthuridae?

References:

Bailly, N., 2010. Why there may be discrepancies in the assessment of scientific names between the Catalog of Fishes and FishBase. Electronic document. [Version 2, May 6th, 2010]. //www.fishbase.org/Nomenclature/FBCofFNames.php

Eschmeyer, W.N. (ed.). Catalog of Fishes. Successive updated online versions http://research.calacademy.org/research/Ichthyology/catalog/fishcatmain.asp.

Eschmeyer, W.N. and J.D. Fong. Species of Fishes by family/subfamily. Successive updated online versions. http://research.calacademy.org/research/ichthyology/catalog/SpeciesByFamily.asp.

Minelli, A., 2009. Perspectives in animal phylogeny. Oxford University Press.

Nelson, J.S., 2006. Fishes of the World. 4th ed. Hoboken (New Jersey, USA): John Wiley & Sons. xix+601 p. [Ref. 58010]

Wiley, E.O. and G.D. Johnson, 2010. pp. 123-182. A teleost classification based on monophyletic groups. In: Origin and phylogenetic interrelationships of Teleosts. Honoring Gloria Arratia. Proceedings of the International Symposium at the ASIH Annual Meeting in St. Louis, Missouri, 2007. (Nelson et al., eds). München (Germany): Verlag Dr. Friedrich Pfeil. [Ref. 84927]

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