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<taxon>
	<dc:identifier>FB-106</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=106</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Scombriformes</dwc:Order>
	<dwc:Family>Scombridae</dwc:Family>
	<dwc:Genus>Gymnosarda</dwc:Genus>
	<dwc:ScientificName>Gymnosarda unicolor (Rüppell, 1836)</dwc:ScientificName>
	<commonName xml:lang='fa'>Posht Siah</commonName>
	<commonName xml:lang='fa'>Puzanok</commonName>
	<commonName xml:lang='fa'>Zalun</commonName>
	<commonName xml:lang='pl'>Sledz czarnomorski</commonName>
	<commonName xml:lang='pt'>Sável-do-Mar Negro</commonName>
	<commonName xml:lang='bg'>Dunavska skumriya</commonName>
	<commonName xml:lang='bg'>Karagyoz</commonName>
	<commonName xml:lang='da'>Eichwalds stamsild</commonName>
	<commonName xml:lang='de'>Donauhering</commonName>
	<commonName xml:lang='en'>Black Sea herring</commonName>
	<commonName xml:lang='en'>Black Sea shad</commonName>
	<commonName xml:lang='en'>Kerch black sea shad</commonName>
	<commonName xml:lang='en'>Kerch shad</commonName>
	<commonName xml:lang='en'>Pontic shad</commonName>
	<commonName xml:lang='en'>Spotless shad</commonName>
	<commonName xml:lang='et'>Musta ja aasovi mere aloosa</commonName>
	<commonName xml:lang='et'>Pontose aloosa</commonName>
	<commonName xml:lang='fr'>Alose</commonName>
	<commonName xml:lang='fr'>Alose de la mer Noire</commonName>
	<commonName xml:lang='ru'>Chernomorskaya seld</commonName>
	<commonName xml:lang='ru'>Chernomorskaya sel'd</commonName>
	<commonName xml:lang='ru'>Chernomorsko-azovskaya sel'd'</commonName>
	<commonName xml:lang='ru'>Dunayskaya sel'd</commonName>
	<commonName xml:lang='ru'>Tchernomorsko-asovskaya sel'd</commonName>
	<commonName xml:lang='ru'>Сельдь черноморско-азовская</commonName>
	<commonName xml:lang='ru'>черноморско-азовская проходная сельдь</commonName>
	<commonName xml:lang='sv'>Svartahavs-staksill</commonName>
	<commonName xml:lang='tr'>Ringa balığı</commonName>
	<commonName xml:lang='tr'>Tirsi</commonName>
	<commonName xml:lang='tr'>Tirsi balığı</commonName>
	<commonName xml:lang='uk'>Оселедець азово-чорноморський</commonName>
	<commonName xml:lang='uk'>Оселедець чорноморський</commonName>
	<commonName xml:lang='uk'>Оселедець чорноморсько-азовський</commonName>
	<commonName xml:lang='uk'>Оселедець­куцак</commonName>
	<commonName xml:lang='es'>Alosa pontica</commonName>
	<commonName xml:lang='es'>Sábalo del Mar Negro</commonName>
	<commonName xml:lang=''>Črnomorska čepa</commonName>
	<commonName xml:lang=''>Sardelomana Pontou</commonName>
	<commonName xml:lang=''>Scrumbie de Dunare</commonName>
	<commonName xml:lang=''>Scumbie de Dunăre</commonName>
	<commonName xml:lang=''>Σαρδελομάνα  Πόντου</commonName>
	<commonName xml:lang=''>長體西鯡</commonName>
	<commonName xml:lang=''>长体西鲱</commonName>
	<synonym relationship='synonym'>Alosa kessleri pontica (Eichwald, 1838)</synonym>
	<synonym relationship='ambiguous synonym'>Alosa kessleri pontica borysthenis Pavlov, 1959</synonym>
	<synonym relationship='ambiguous synonym'>Alosa kessleri pontica issatschenkovi Pavlov, 1959</synonym>
	<synonym relationship='ambiguous synonym'>Alosa kessleri pontica moriaac Pavlov, 1959</synonym>
	<synonym relationship='synonym'>Alosa pontica (Eichwald, 1838)</synonym>
	<synonym relationship='ambiguous synonym'>Alosa pontica danubii Antipa, 1904</synonym>
	<synonym relationship='ambiguous synonym'>Alosa pontica nigrescens Antipa, 1904</synonym>
	<synonym relationship='synonym'>Alosa pontica pontica (Eichwald, 1838)</synonym>
	<synonym relationship='ambiguous synonym'>Alosa pontica russac Antipa, 1904</synonym>
	<synonym relationship='synonym'>Alosa pontica var. moriac Antipa, 1909</synonym>
	<synonym relationship='synonym'>Caspialosa kessleri pontica (Eichwald, 1838)</synonym>
	<synonym relationship='synonym'>Caspialosa pontica (Eichwald, 1838)</synonym>
	<synonym relationship='ambiguous synonym'>Caspialosa pontica chtamalocephala Isachenko, 1925</synonym>
	<synonym relationship='ambiguous synonym'>Caspialosa pontica hypselocephala Isachenko, 1925</synonym>
	<synonym relationship='synonym'>Clupea eichwaldi Grimm, 1901</synonym>
	<synonym relationship='synonym'>Clupea eichwaldii Grimm, 1901</synonym>
	<synonym relationship='misapplied name'>Clupea piltschardus (non Pallas, 1814)</synonym>
	<synonym relationship='synonym'>Clupea pontica Eichwald, 1838</synonym>
	<dcterms:created>1990-10-17</dcterms:created>
	<dcterms:modified>2023-06-15</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96734&amp;speccode=105'>Ergudren, E., F. Turan and C. Turan0 Length-weight and length-length relationships for four shad species along the western Black Sea coast of Turkey. J. Appl. Ichthyol. 27:942-944. (Ref. 96734)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=105'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=105'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105574&amp;speccode=105'>Termeki on Eesti Keele Instituudi0 Termeki on Eesti Keele Instituudi. http://term.eki.ee/termbase/view/7351963/ (Ref. 105574)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106151&amp;speccode=105'>Çiçek, E., R. Fricke and S.S. Birecikligil0 Freshwater fishes of Turkey: a revised and updated annotated checklist. Biharean Biologist 9(2):141-157. (Ref. 106151)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=107945&amp;speccode=105'>Cheung, W.L., R. Watson and D. Pauly0 Signature of ocean warming in global fisheries catch. Nature 497:365-368. (Ref. 107945)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114869&amp;speccode=105'>Ibanescu, D.C., A. Popescu and A. Nica0 Estimation of growth and mortalty parameters of the Pontic shad (&lt;i&gt;Alosa immaculata&lt;/i&gt; Bennett, 1835) in Romanian section of the Danube River. Lucrări Ştiinţifice-Seria Zootehnie: 285-289. https://www.researchgate.net/publication/311651581 (Ref. 114869)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=105'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115544&amp;speccode=105'>Keskin, C.0 A preliminary study on demersal fishes in the southwestern Black Sea shelf (NW Turkey). J. Black Sea/Mediterranean Environment 18(3):341-349. (Ref. 115544)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=105'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=105'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121643&amp;speccode=105'>Samsun, O., O. Akyol, T. Ceyhan and Y. Erden0 Length-weight relationships for 11 fish species from the Central Black Sea, Turkey. Ege Journal of Fisheries and Aquatic Sciences 34(4):455-458. (Ref. 121643)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124004&amp;speccode=105'>Kuljanishvili, T., G. Epitashvili, J. Freyhof, B. Japoshvili, L. Kalous, B. Levin, N. Mustafayev, S. Ibrahimov, S. Pipoyan and L. Mumladze0 Checklist of the freshwater fishes of Armenia, Azerbaijan and Georgia. J. Appl. Ichthyol. 36(4):501-514. (Ref. 124004)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=105'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=105'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=105'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard0 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference><dataObject>
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	<dc:description>Thai species is pelagic at sea, in deep water.  It migrates to middle reaches of large rivers, spawning where current is strongest, close to the surface, often at 2-3 m depth in the main channel.  Migrates upriver to spawn at 3 years, rarely earlier and only a few individuals spawn two seasons.  It appears along the coast in March to April, enters rivers when temperatures reach about 6-9°C, between late March and late April; migration usually peaks in May.  Spawning starts when temperature rises above 15°C in April to August; usually between 1 and 8 p.m.  Pelagic eggs.  Spent individuals return to the sea to feed.  Juveniles inhabit floodplain and shallow riverine habitats, migrate to the sea or estuarine habitats during first summer; in autumn return to the sea until maturity.  When at sea, it feeds on a wide variety of zooplankton (mainly crustaceans, (Crangon, Upogebia, Idothea, gammarids) and small fish (Engraulis, Clupeonella, Sprattus).  Reduction of spawning sites and migration routes are caused by impoundment of main rivers.  Heavy over fishing apparently reduced all populations during the first decade of the 20th century (Ref. 59043).  Reaches a smaller size (30 cm SL, usually 14-18) in Don River; a smaller form reaches only 21 cm.</dc:description>
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	<dc:description>Body fairly elongate, more `herring-like' than `shad-like'.  Gill rakers rather thin, usually equal to or a little shorter than gill filaments.  Teeth well developed in both jaws.  Resembles A. caspia, which usually has more Gill rakers (50 to 180, much longer than gill filaments), poorly developed teeth and a deeper, `shad-like' body; A. maeotica has fewer Gill rakers (33 to 36).</dc:description>
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	<dc:description>Dorsal spines (total): 0;  Analspines: 0</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=188&amp;speccode=105'>Whitehead, P.J.P. 1985 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188)</reference></dataObject><dataObject>
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	<dc:description>Eurasia:  Black Sea and Sea of Azov ( in sea and in the Don, Danube and other rivers, as much as 567 km up the Don and as far as Kiev on the Dneiper before the dam was built).  Recognized sub-species (Ref. 683):  Alosa pontica pontica in the Black Sea and rivers feeding it; Alosa pontica kessleri and Alosa pontica volgensis in the Caspian Sea.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26334&amp;speccode=105'>Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov 1997 An annotated check-list of the freshwater fishes of Russia. J. Ichthyol. 37(9):687-736. (Ref. 26334)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4931&amp;speccode=105'>FAO 1992 FAO yearbook. Fishery statistics: catches and landings. Vol. 70: 1990. FAO Fish. Ser. 38:647 p. (Ref. 4931)</reference></dataObject><dataObject>
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	<dc:description>Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous.    Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=105'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard 2009 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference></dataObject><dataObject>
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	<dc:description>Eggs are pelagic (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=188&amp;speccode=105'>Whitehead, P.J.P. 1985 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188)</reference></dataObject><dataObject>
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	<dc:description>Feeds on fish and benthic crustaceans (Ref. 188). Euryhaline, anadromous, moving northward in the Black Sea in spring and early summer. Remains at depths of 40-90 m during winter, moving to shallower waters in spring and near the bottom  at water temperatures of 10-15°C in summer and autumn (Ref. 10439). In Kagoul Lake in the Danube Delta, numerous juveniles 2.6-7.0 cm FL were noted in July-August at water temperatures of 15.8-19.5°C, oxygen concentration of 9.9-10.4 ml/l and pH of 8.0-8.1 (Ref. 10440).</dc:description>
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	<dc:description>39.0 cm SL (male/unsexed; (Ref. 188)); max. reported age: 7 years (Ref. 10439)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=188&amp;speccode=105'>Whitehead, P.J.P. 1985 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10439&amp;speccode=105'>Kolarov, P. 1991 &lt;i&gt;Alosa pontica pontica&lt;/i&gt; (Eichwald 1938). p. 337-368. In H. Hoestlandt (ed.) The freshwater fishes of Europe. Vol. 2. Clupeidae, Anguillidae. AULA-Verlag Wiesbaden. (Ref. 10439)</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject></subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-105</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1990-10-17 00:00:00</dcterms:created>
	<dcterms:modified>2025-10-21 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder> FAO</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=105</dc:source>
	<subject></subject>
	<dc:description>pelagic-neritic; anadromous (Ref. 138126); freshwater; brackish; marine; depth range 3 - 90 m (Ref. 10439)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138126&amp;speccode=105'>Lassalle, G., M. Béguer, L. Baulaton and E. Rochard 2009 Learning from the past to predict the future: Responses of European diadromous fish to climate change. Amer. Fish. Soc. Symp. 69:175-193. (Ref. 138126)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10439&amp;speccode=105'>Kolarov, P. 1991 &lt;i&gt;Alosa pontica pontica&lt;/i&gt; (Eichwald 1938). p. 337-368. In H. Hoestlandt (ed.) The freshwater fishes of Europe. Vol. 2. Clupeidae, Anguillidae. AULA-Verlag Wiesbaden. (Ref. 10439)</reference></dataObject>
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