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<taxon>
	<dc:identifier>FB-3213</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3213</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cyprinodontiformes</dwc:Order>
	<dwc:Family>Rivulidae</dwc:Family>
	<dwc:Genus>Kryptolebias</dwc:Genus>
	<dwc:ScientificName>Kryptolebias marmoratus (Poey, 1880)</dwc:ScientificName>
	<commonName xml:lang='de'>Marmorierter Bachling</commonName>
	<commonName xml:lang='en'>Mangrove rivulus</commonName>
	<commonName xml:lang='en'>Mangrove Rivulus</commonName>
	<commonName xml:lang='en'>Matanzas Rivulus</commonName>
	<commonName xml:lang='en'>Matanzas rivulus</commonName>
	<commonName xml:lang='en'>Rivulus</commonName>
	<commonName xml:lang='es'>Almirante de manglar</commonName>
	<commonName xml:lang='es'>Rivulo De Mangle</commonName>
	<commonName xml:lang='es'>Rivulo Matancero</commonName>
	<commonName xml:lang=''>Kamulibo</commonName>
	<commonName xml:lang=''>Loangouma</commonName>
	<commonName xml:lang=''>Toumblouc</commonName>
	<commonName xml:lang=''>Wilili</commonName>
	<commonName xml:lang=''>加尔西氏溪鮰</commonName>
	<commonName xml:lang=''>加爾西氏溪鮰</commonName>
	<commonName xml:lang=''>花斑溪鮰</commonName>
	<synonym relationship='synonym'>Rivulus bonairensis Hoedeman, 1958</synonym>
	<synonym relationship='synonym'>Rivulus garciae de la Cruz &amp; Dubitsky, 1976</synonym>
	<synonym relationship='synonym'>Rivulus garciai de la Cruz &amp; Dubitsky, 1976</synonym>
	<synonym relationship='synonym'>Rivulus heyei Nichols, 1914</synonym>
	<synonym relationship='synonym'>Rivulus marmoratus Poey, 1880</synonym>
	<synonym relationship='synonym'>Rivulus marmoratus bonairensis Hoedeman, 1958</synonym>
	<synonym relationship='misapplied name'>Rivulus occellatus (non Hensel, 1868)</synonym>
	<synonym relationship='misapplied name'>Rivulus ocellatus (non Hensel, 1868)</synonym>
	<dcterms:created>1991-05-01</dcterms:created>
	<dcterms:modified>2024-01-15</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=63&amp;speccode=3213'>ASIH (American Society of Ichthyologists and Herpetologists)0 Ontogeny and systematics of fishes. Based on an international symposium dedicated to the memory of Elbert Halvor Ahlstrom, 15-18 August 1983, La Jolla, California. Spec. Publ. Am. Soc. Ichthyol. Herpetol. 1:ix, 760 p. (Ref. 63)</reference>
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	<reference isbn='2-86515-117-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=35237&amp;speccode=3213'>Keith, P., P.-Y. Le Bail and P. Planquette0 Atlas des poissons d'eau douce de Guyane. Tome 2, Fascicule I: Batrachoidiformes, Mugiliformes, Beloniformes, Cyprinodontiformes, Synbranchiformes, Perciformes, Pleuronectiformes, Tetraodontiformes. Collection Patrimoines Naturels 43(I): 286p. Paris: Publications scientifiques du Muséum national d'Histoire naturelle. (Ref. 35237)</reference>
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	<reference isbn='92-5-104826-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=50882&amp;speccode=3213'>Ghedotti, M.J. and E.O. Wiley0 Rivulidae. New World rivulines (rivulid killifishes). p. 1145-1146. In K.E. Carpenter (ed.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Atlantic. Vol. 2: Bony fishes part 1 (Acipenseridae to Grammatidae). (Ref. 50882)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52570&amp;speccode=3213'>Costa, W.J.E.M.0 A new killifish genus and species from the coastal plains of north-eastern Brazil (Teleostei: Cyprinodontiformes: Rivulidae). Zootaxa 642(1):1-10. (Ref. 52570)</reference>
	<reference isbn='1-56098-985-8' url='http://www.fishbase.org/references/FBRefSummary.php?id=54206&amp;speccode=3213'>Claro, R. and L.R. Parenti0 The marine ichthyofauna of Cuba. p. 21-57. In Claro, R., K.C. Lindeman and L.R. Parenti (eds) Ecology of the marine fishes of Cuba. Smithsonian Institution Press, Wahsington and London. 253p. (Ref. 54206)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55170&amp;speccode=3213'>Florida Museum of Natural History0 Biological profiles: mangrove rivulus. Retrieved on 26 August 2005, from www.flmnh.ufl.edu/fish/Gallery/Descript/MangroveRivulus/MangroveRivulus.html. Ichthyology at the Florida Museum of Natural History: Education-Biological Profiles. FLMNH, University of Florida. (Ref. 55170)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=3213'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80809&amp;speccode=3213'>Costa, W.J.E.M.0 Redescription of &lt;i&gt;Kryptolebias ocellatus&lt;/i&gt; (Hensel) and &lt;i&gt;K. caudomarginatus&lt;/i&gt; (Seegers) (Teleostei: Cyprinodontiformes: Rivulidae), two killifishes from mangroves of south-eastern Brazil. aqua, J. Ichthyol. Aquat. Biol. 11(1):5-12. (Ref. 80809)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81264&amp;speccode=3213'>Jelks, H.L., S.J. Walsh, N.M. Burkhead, S. Contreras-Balderas, E. Díaz-Pardo, D.A. Hendrickson, J. Lyons, N.E. Mandrak, F. McCormick, J.S. Nelson, S.P. Platania, B.A. Porter, C.B. Renaud, J.J. Schmitter-Soto, E.B. Taylor and M.L. Warren Jr.0 Conservation status of imperiled North American freshwater and diadromous fishes. Fisheries 33(8):372-407. (Ref. 81264)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93034&amp;speccode=3213'>Edenbrow, M. and D.P. Croft0 Behavioural types and life history strategies during ontogeny in the mangrove killifish, &lt;i&gt;Kyrptolebias marmoratus&lt;/i&gt;. Anim. Behav. 82(4):731-741. (Ref. 93034)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126106&amp;speccode=3213'>Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante0 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=3213'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-12-12 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
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	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=3213</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Adults inhabit shallow, mud-bottomed ditches, bays, salt marshes and other brackish-water environments; also crab burrows (Ref. 5723), moist leaf litter, decaying mangrove logs (Ref. 93034).  Can withstand heavy pollution of fresh or salty waters and shares its biotope with Poecilia vivipara which is much more abundant (Ref. 35237).  Usually found in water with low oxygen content (Ref. 7251).  Either male or hermaphroditic, females don't seem to exist.  Only about 5% of a population are born as males; after 3-4 years about 60% of the (self-fertilizing) hermaphrodites transform into secondary males by losing female structure and function.  The proportion of males depends on the environmental temperature.  Below 20°C, majority are males, above 25°C all are hermaphrodites (Ref. 35237).  It is the only known naturally occurring, self-fertilizing vertebrate (Ref. 13056, 35237).  Not a seasonal killifish.  Is very difficult to maintain in aquarium (Ref. 27139).</dc:description>
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	<reference isbn='85-7430-361-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=36579&amp;speccode=3213'>Costa, W.J.E.M. 2003 Rivulidae (South American Annual Fishes). p. 526-548. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil. (Ref. 36579)</reference></dataObject><dataObject>
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	<dc:description>Dark spot surrounded by yellow ring on side of caudal peduncle, just ahead of upper part of caudal fin base (Ref. 26938).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26938&amp;speccode=3213'>Smith, C.L. 1997 National Audubon Society field guide to tropical marine fishes of the Caribbean, the Gulf of Mexico, Florida, the Bahamas, and Bermuda. Alfred A. Knopf, Inc., New York. 720 p. (Ref. 26938)</reference></dataObject><dataObject>
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	<dc:description>North, Central and South America:  eastern coasts of Florida (Indian River to Key West) in USA and Bahamas to Cuba, Jamaica, Cayman Island, Brazil, French Guiana, the Netherlands Antilles, Venezuela, Belize, Mexico, Puerto Rico.  Throughout Caribbean (Ref. 26938).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27139&amp;speccode=3213'>Huber, J.H. 1996 Killi-Data 1996. Updated checklist of taxonomic names, collecting localities and bibliographic references of oviparous Cyprinodont fishes (Atherinomorpha, Pisces). Société Française d'Ichtyologie, Muséum National d'Histoire Naturelle, Paris, France, 399 p. (Ref. 27139)</reference></dataObject><dataObject>
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	<dcterms:modified>2025-12-12 00:00:00</dcterms:modified>
	<dc:title>Importance</dc:title>
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	<dc:description>fisheries: commercial; aquarium: commercial; bait:  </dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27139&amp;speccode=3213'>Huber, J.H. 1996 Killi-Data 1996. Updated checklist of taxonomic names, collecting localities and bibliographic references of oviparous Cyprinodont fishes (Atherinomorpha, Pisces). Société Française d'Ichtyologie, Muséum National d'Histoire Naturelle, Paris, France, 399 p. (Ref. 27139)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27139&amp;speccode=3213'>Huber, J.H. 1996 Killi-Data 1996. Updated checklist of taxonomic names, collecting localities and bibliographic references of oviparous Cyprinodont fishes (Atherinomorpha, Pisces). Société Française d'Ichtyologie, Muséum National d'Histoire Naturelle, Paris, France, 399 p. (Ref. 27139)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=3213&amp;StockCode=3409</dc:source>
	<dc:description>Eggs are sometimes fertilized internally by accident via close apposition of gonopores in otherwise oviparous fishes (Ref. 7471). Regarded as a self-fertilizing hermaphrodite (Ref. 6028, 245, 35237). Eggs are 1.6 mm in diameter. Sexes can be distinguished after 8 weeks (Ref. 35237).  Also Ref. 103751.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7471&amp;speccode=3213'>Balon, E.K. 1990 Epigenesis of an epigeneticist: the development of some alternative concepts on the early ontogeny and evolution of fishes. Guelph Ichthyol. Rev. 1:1-48. (Ref. 7471)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-3213</dc:identifier>
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	<dc:title>Max. size</dc:title>
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	<dc:description>7.5 cm TL (male/unsexed; (Ref. 27139))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27139&amp;speccode=3213'>Huber, J.H. 1996 Killi-Data 1996. Updated checklist of taxonomic names, collecting localities and bibliographic references of oviparous Cyprinodont fishes (Atherinomorpha, Pisces). Société Française d'Ichtyologie, Muséum National d'Histoire Naturelle, Paris, France, 399 p. (Ref. 27139)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-3213</dc:identifier>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-05-01 00:00:00</dcterms:created>
	<dcterms:modified>2025-12-12 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>benthopelagic; non-migratory (Ref. 126106); freshwater; brackish; pH range: 7.5; dH range: 16; depth range 0 - ? m (Ref. 55170)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126106&amp;speccode=3213'>Dean, E.M., A.R. Cooper, L. Wang, W. Daniel, S. David, C. Ernzen, K.B. Gido, E Hale, T.J. Haxton, W. Kelso, N. Leonard, C. Lido, J. Margraf, M. Porter, C. Pennock, D. Propst, J. Ross, M.D. Staudinger, G. Whelan and D.M. Infante 2022 The North American Freshwater Migratory Fish Database (NAFMFD): Characterizing the migratory life histories of freshwater fishes of Canada, the United States of Mexico. J. Biogeography 49:1193-1203. (Ref. 126106)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55170&amp;speccode=3213'>Florida Museum of Natural History 2005 Biological profiles: mangrove rivulus. Retrieved on 26 August 2005, from www.flmnh.ufl.edu/fish/Gallery/Descript/MangroveRivulus/MangroveRivulus.html. Ichthyology at the Florida Museum of Natural History: Education-Biological Profiles. FLMNH, University of Florida. (Ref. 55170)</reference></dataObject>
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