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	<dc:identifier>FB-4503</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Anabantiformes</dwc:Order>
	<dwc:Family>Osphronemidae</dwc:Family>
	<dwc:Genus>Luciocephalus</dwc:Genus>
	<dwc:ScientificName>Luciocephalus pulcher (Gray, 1830)</dwc:ScientificName>
	<commonName xml:lang='pl'>Szczupakoglów</commonName>
	<commonName xml:lang='da'>Geddehoved</commonName>
	<commonName xml:lang='da'>Geddehovedfisk</commonName>
	<commonName xml:lang='de'>Hechtkopf</commonName>
	<commonName xml:lang='en'>Pikehead</commonName>
	<commonName xml:lang='fi'>Hauenpääkala</commonName>
	<commonName xml:lang='ms'>Emplunyu</commonName>
	<commonName xml:lang='ms'>Plunyu</commonName>
	<commonName xml:lang='ms'>Tembok tebing</commonName>
	<commonName xml:lang='ru'>щукоглав</commonName>
	<commonName xml:lang='sv'>Gäddhuvudfisk</commonName>
	<commonName xml:lang=''>梭头鲈</commonName>
	<commonName xml:lang=''>梭頭鱸</commonName>
	<synonym relationship='synonym'>Diplopterus pulcher Gray, 1830</synonym>
	<dcterms:created>1991-06-28</dcterms:created>
	<dcterms:modified>2008-02-06</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=245&amp;speccode=4503'>Nelson, J.S.0 Fishes of the world. 2nd edition. John Wiley &amp; Sons, Inc., New York. 523 p. (Ref. 245)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1672&amp;speccode=4503'>Riehl, R. and H.A. Baensch0 Aquarien Atlas. Band. 1. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 992 p. (Ref. 1672)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=4503'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4835&amp;speccode=4503'>Mohsin, A.K.M. and M.A. Ambak0 Freshwater fishes of Peninsular Malaysia. Penerbit Universiti Pertanian Malaysia, Malaysia. 284 p. (Ref. 4835)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7050&amp;speccode=4503'>Kottelat, M., A.J. Whitten, S.N. Kartikasari and S. Wirjoatmodjo0 Freshwater fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong. 221 p. (Ref. 7050)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10937&amp;speccode=4503'>Anonymous0 Fish collection database of the University of British Columbia Fish Museum Fish Museum. University of British Columbia, Vancouver, Canada. (Ref. 10937)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12965&amp;speccode=4503'>Eschmeyer, W.N.0 PISCES. Eschmeyer's PISCES database as published on the Internet in June 1997, URL: gopher://gopher.calacademy.org:640/7. (Ref. 12965)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13371&amp;speccode=4503'>Riehl, R. and H.A. Baensch0 Aquarien Atlas, Band 1. 10th edition. Mergus Verlag GmBH, Melle, Germany. 992 p. (Ref. 13371)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=4503'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26492&amp;speccode=4503'>Kokoscha, M.0 Fortpflanzung bei der Gattung &lt;i&gt;Luciocephalus&lt;/i&gt;. p. 33-36. In H. Greven and R. Riehl (eds.) Fortpflanzungsbiologie der Aquarienfische. Birgit Schmettkamp Verlag, Bornheim, Germany. (Ref. 26492)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=4503'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=4503'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=4503'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='974-89954-6-1.' url='http://www.fishbase.org/references/FBRefSummary.php?id=37773&amp;speccode=4503'>Monkolprasit, S., S. Sontirat, S. Vimollohakarn and T. Songsirikul0 Checklist of Fishes in Thailand. Office of Environmental Policy and Planning, Bangkok, Thailand. 353 p. (Ref. 37773)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=4503'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=4503'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=4503'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47438&amp;speccode=4503'>Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48321&amp;speccode=4503'>Ricker, W.E.0 Russian-English dictionary for students of fisheries and aquatic biology. Fisheries Research Board of Canada, Ottawa. (Ref. 48321)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=4503'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=4503'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng0 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56749&amp;speccode=4503'>Kottelat, M. and E. Widjanarti0 The fishes of Danau Sentarum National Park and the Kapuas Lakes area, Kalimantan Barat, Indonesia. Raffles Bull. Zool. Supplement (13):139-173. (Ref. 56749)</reference>
	<reference isbn='974087704-3-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=57235&amp;speccode=4503'>Vidthayanon, C.0 Peat swamp fishes of Thailand. Office of Environmental Policy and Planning, Bangkok, Thailand, 136 p. (Ref. 57235)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57780&amp;speccode=4503'>Parenti, L.R. and K.K.P. Lim0 Fishes of the Rajang basin, Sarawak, Malaysia. Raffles Bull. Zool. Supplement (13):175-208. (Ref. 57780)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57864&amp;speccode=4503'>BFAR0 List of ornamental fish species introduced to the Philippines through NAIA. Unpublished. (Ref. 57864)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=4503'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=85309&amp;speccode=4503'>Lim, K.K.P., H.H. Tan and J.K.Y. Low0 Fishes. p. 145-154. In G.W.H. Davison, P.KL. Ng and H.H. Chew (eds.). The Singapore red data book: threatened plants &amp; animals of Singapore. 2nd ed. Nature Society (Singapore), Singapore. viii, 285 p. : col. ill.; 26 cm. (Ref. 85309)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=85350&amp;speccode=4503'>Lim, K.K.P.0 Checklists of threatened species: fishes, amphibians and reptiles. p. 263-264. In G.W.H. Davison, P.KL. Ng and H.H. Chew (eds.). The Singapore red data book: threatened plants &amp; animals of Singapore. 2nd ed. Nature Society (Singapore), Singapore. viii, 285 p. : col. ill.; 26 cm. (Ref. 85350)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=4503'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-4503</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-28 00:00:00</dcterms:created>
	<dcterms:modified>2008-02-06 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Occurs in streams and flooded forest among dense vegetation (Ref. 56749).  Found in  primary peats swamps.  Mouth brooder (Ref. 57235).  Highly protrusible jaws.  The premaxillae can extend about 1/3 of the head length.  Prey is captured by the fish making a rapid lunge and surrounding the prey with the open mouth.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=245&amp;speccode=4503'>Nelson, J.S. 1984 Fishes of the world. 2nd edition. John Wiley &amp; Sons, Inc., New York. 523 p. (Ref. 245)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-4503</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2007-11-17 00:00:00</dcterms:created>
	<dcterms:modified>2007-11-17 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Distinguished from Luciocephalus aura in the following characters: absence of numerous iridescent green spots on central stripe when live (vs. presence); presence of black irregular pattern on area below preorbital to central stripe (vs. absence); area just above anal-fin base with 3-4 dark brown blotches, which appear circular when body is viewed ventrally (vs. absence of such markings); yellowish caudal fin with 3-5 broad vertical black bars (vs. brownish caudal fin with 3-7 narrow iridescent vertical gold bars); presence of markings on anal and pelvic (vs. absence); shorter caudal-fin length (total length 120.0-125.6% SL vs. 123.7-127.1); shorter postdorsal length (10.4-12.4% SL vs. 12.0-13.2); greater head length (44.1-48.6% SL vs. 43.3-45.9); smaller caudal peduncle depth (8.9-10.6% SL vs 10.7-11.2); and relatively shorter anal-fin base length (19.3-23.4% SL vs. 23.6-24.5) (Ref. 56386).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng 2005 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-4503</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Dorsal soft rays (total): 10 - 12; Anal spines: 0 - 1; Analsoft rays: 19 - 21; Vertebrae: 38 - 40</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng 2005 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Kent E. Carpenter</agent>
	<dcterms:created>1992-06-15 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Asia:  Peninsular Thailand, Malaysia, Brunei Darussalam and Indonesia (Ref. 56386) and Singapore (Ref. 85309).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng 2005 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-4503</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-28 00:00:00</dcterms:created>
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	<dc:title>Importance</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<dc:description>fisheries: commercial; aquarium: commercial</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng 2005 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56386&amp;speccode=4503'>Tan, H.H. and P.K.L. Ng 2005 The labyrinth fishes (Teleostei: Anabantoidei, Channoidei) of Sumatra, Indonesia. Raffles Bull. Zool. Supplement (13):115-138. (Ref. 56386)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-4503</dc:identifier>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1991-06-28 00:00:00</dcterms:created>
	<dcterms:modified>2008-02-06 00:00:00</dcterms:modified>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=4503&amp;StockCode=6143</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Males are mouthbrooding for about 30 days.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26492&amp;speccode=4503'>Kokoscha, M. 1995 Fortpflanzung bei der Gattung &lt;i&gt;Luciocephalus&lt;/i&gt;. p. 33-36. In H. Greven and R. Riehl (eds.) Fortpflanzungsbiologie der Aquarienfische. Birgit Schmettkamp Verlag, Bornheim, Germany. (Ref. 26492)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-4503</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1991-06-28 00:00:00</dcterms:created>
	<dcterms:modified>2008-02-06 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>20.0 cm SL (male/unsexed; (Ref. 57235))</dc:description>
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	<reference isbn='974087704-3-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=57235&amp;speccode=4503'>Vidthayanon, C. 2002 Peat swamp fishes of Thailand. Office of Environmental Policy and Planning, Bangkok, Thailand, 136 p. (Ref. 57235)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=4503</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>benthopelagic; freshwater; pH range: 6.0 - 7.5; dH range: 15</dc:description>
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