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	<dc:identifier>FB-51557</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=51557</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Blenniiformes</dwc:Order>
	<dwc:Family>Tripterygiidae</dwc:Family>
	<dwc:Genus>Enneapterygius</dwc:Genus>
	<dwc:ScientificName>Enneapterygius signicauda Fricke, 1997</dwc:ScientificName>
	<commonName xml:lang='en'>Triangle triplefin</commonName>
	<commonName xml:lang='ja'>Aya-hebiginpo</commonName>
	<commonName xml:lang=''>哈氏弯线鳚</commonName>
	<commonName xml:lang=''>哈氏彎線鳚</commonName>
	<commonName xml:lang=''>屈体三鳍鳚</commonName>
	<commonName xml:lang=''>屈體三鰭鳚</commonName>
	<commonName xml:lang=''>線頭彎線鳚</commonName>
	<commonName xml:lang=''>线头弯线鳚</commonName>
	<synonym relationship='synonym'>Enneapterygius inclinatus Fowler, 1946</synonym>
	<synonym relationship='synonym'>Helcogramma habena Williams &amp; McCormick, 1990</synonym>
	<synonym relationship='misapplied name'>Helcogramma hudsoni (non Jordan &amp; Seale, 1906)</synonym>
	<synonym relationship='synonym'>Helcogramma inclinatum (Fowler, 1946)</synonym>
	<synonym relationship='synonym'>Tripterygion inclinatum (Fowler, 1946)</synonym>
	<dcterms:created>1999-02-19</dcterms:created>
	<dcterms:modified>2021-10-06</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=240&amp;speccode=51565'>Thresher, R.E.0 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9849&amp;speccode=51565'>Williams, J.T. and C.J. McCormick0 Two new species of the triplefin fish genus &lt;i&gt;Helcogramma&lt;/i&gt; (Tripterygiidae) from the Western Pacific Ocean. Copeia 1990(4):1020-1030. (Ref. 9849)</reference>
	<reference isbn='3-87429-400-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27223&amp;speccode=51565'>Fricke, R.0 Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=51565'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31667&amp;speccode=51565'>Anonymous0 Fish collection database of the Bernice P. Bishop Museum (BPBM). Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36648&amp;speccode=51565'>Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=51565'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47843&amp;speccode=51565'>Huang, Z.0 Marine species and their distribution in China's seas. p. 404-463. Vertebrata. Smithsonian Institution, Florida, USA. 598 p. (Ref. 47843)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52308&amp;speccode=51565'>Williams, J.T. and J.C. Howe0 Seven new species of the triplefin fish genus &lt;i&gt;Helcogramma&lt;/i&gt; (Tripterygiidae) from the Indo-Pacific. aqua, J. Ichthyol. Aquat. Biol. 7(4):151-176. (Ref. 52308)</reference>
	<reference isbn='957-01-6879-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=55073&amp;speccode=51565'>Chen, C.-H.0 Checklist of the fishes of Penghu. FRI Special Publication No. 4. 175 p. (Ref. 55073)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58018&amp;speccode=51565'>Bogutskaya, N.G.0 Preliminary assignment of coordinates to type localities in the Catalog of Fishes. Unpublished dbf file. (Ref. 58018)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=51565'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=51565'>Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93609&amp;speccode=51565'>Patzner, R.A., E.J. Gonçalves, P.A. Hastings and B.G. Kapoor0 The biology of blennies. Enfield, NH : Science Publishers, 482 p. (Ref. 93609)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94114&amp;speccode=51565'>Watson, W.0 Larval development in blennies. pp. 309-350. In Patzner, R.A., E.J. Gonçalves, P.A. Hastings and B.G. Kapoor (eds.) The biology of blennies. Science Publishers, Enfield, NH, USA. 482 p. (Ref. 94114)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94457&amp;speccode=51565'>Chiang, M.-C. and I.-S. Chen0 A new species of the genus &lt;i&gt;Helcogramma&lt;/i&gt; (Blenniiformes, Tripterygiidae) from Taiwan. ZooKeys 216: 57-72. (Ref. 94457)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114924&amp;speccode=51565'>Fricke, R. and M.V. Erdmann0 &lt;i&gt;Helcogramma atauroensis&lt;/i&gt;, a new species of triplefin from Ataúro Island, Timor-Leste, eastern Indian Ocean (Teleostei: Tripterygiidae). J. Ocean Sci. Found. 26:34-45. (Ref. 114924)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-02-19 00:00:00</dcterms:created>
	<dcterms:modified>2014-03-24 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=51565</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>Adults are found in rocky shore areas and rocky surge channels with sparse coral growth at depths of less than a meter to 6 m (Ref. 9849).  Eggs are hemispherical and covered with numerous sticky threads that anchor them in the algae on the nesting sites (Ref. 240).  Larvae are planktonic which occur primarily in shallow, nearshore waters (Ref. 94114).</dc:description>
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	<reference isbn='3-87429-400-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27223&amp;speccode=51565'>Fricke, R. 1997 Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Kristine Elaine Paz Olisa</agent>
	<dcterms:created>2013-06-18 00:00:00</dcterms:created>
	<dcterms:modified>2013-12-04 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Description:  Dorsal rays III + XII -XVI (usually XIV-XV) + 9-11 (usually 11), first dorsal spine remarkably longer than succeeding two, 0-2 pterygiophore without spines; anal fin I, 18-22 (usually 20-21); pectoral rays 16-17 (usually 1+8+7); pelvic fin I,2; caudal rays 13, branched 9, procurrent 8-10 both in dorsal and ventral.  Vertebrae 10+ 27-30; last pleural rib on 10th, last epipleural rib on 13-16th vertebral centrum.  Lateral line continuous, pored scales 24-35 (usually 28); head scaleless, nape 1-2 rows, dorsalmost usually missing; scales ctenoid; largest (twice in size) at the area posterior to pectoral-fin base and below the lateral line.  Gill rakers on first arch 8; pseudobranchial filaments 4; mandibular sensory pores 5-12 + 5-10 + 5-12 (usually 9+5+8); orbital cirrus small (sometimes minute) (Ref. 9849, 52308, 90102).  Overall translucent grey with dark vertical saddles, Y-shaped bars (in females and young males, mainly dorsal) or triangular blotches (in mature males, mainly ventral).  Mature males head darker on ventral side pale on upper half, bicolored pattern extends posteriorly to pectoral-fin area which has 3 pale spots vertically aligned; all other fins uniformly black.  Females and young males, head with prominent blue horizontal stripe below the eye, fins transluscent or pale, dorsal and anal fins with small dark spots, pectoral fin with dark spot over bases of median rays, caudal fin pale with 3 dark stripes (Ref. 9849, 52308, 90102). Also Ref. 94457.</dc:description>
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	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=51565'>Allen, G.R. and M.V. Erdmann 2012 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Kristine Elaine Paz Olisa</agent>
	<dcterms:created>2013-06-18 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=51565</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Dorsal spines (total): 15 - 19; Dorsal soft rays (total): 9 - 11;  Analspines: 1; Analsoft rays: 18 - 22; Vertebrae: 37 - 40</dc:description>
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	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=51565'>Allen, G.R. and M.V. Erdmann 2012 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-02-19 00:00:00</dcterms:created>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Western Pacific:  Ryukyu Islands, Taiwan, and the Philippines.</dc:description>
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	<reference isbn='3-87429-400-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27223&amp;speccode=51565'>Fricke, R. 1997 Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Kristine Elaine Paz Olisa</agent>
	<dcterms:created>2013-06-24 00:00:00</dcterms:created>
	<dcterms:modified>2013-07-05 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
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	<dc:source>inclinata</dc:source>
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	<dc:description>Found in rocky shore areas and rocky surge channels with little coral growth at less than a meter to 6 m deep (Ref. 90102).   Occurs on large encrusted boulders, vertical rock walls, and large shallow caves; surge channel at edge of rock platform, spotty corral, and sand and encrusted rock bottom (Ref. 9849).</dc:description>
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	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=51565'>Allen, G.R. and M.V. Erdmann 2012 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-51565</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-02-19 00:00:00</dcterms:created>
	<dcterms:modified>2014-03-24 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=51565</dc:source>
	<dc:description>4.6 cm SL (male/unsexed; (Ref. 27223))</dc:description>
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	<reference isbn='3-87429-400-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27223&amp;speccode=51565'>Fricke, R. 1997 Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-51565</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1999-02-19 00:00:00</dcterms:created>
	<dcterms:modified>2014-03-24 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=51565</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>demersal; marine; depth range 0 - 9 m (Ref. 27223)</dc:description>
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	<reference isbn='3-87429-400-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=27223&amp;speccode=51565'>Fricke, R. 1997 Tripterygiid fishes of the western and central Pacific, with descriptions of 15 new species, including an annotated checklist of world Tripterygiidae (Teleostei). Theses Zool. 29:1-607. (Ref. 27223)</reference></dataObject>
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