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	<dc:identifier>FB-53919</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=53919</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Torpediniformes</dwc:Order>
	<dwc:Family>Narkidae</dwc:Family>
	<dwc:Genus>Heteronarce</dwc:Genus>
	<dwc:ScientificName>Heteronarce bentuviai (Baranes &amp; Randall, 1989)</dwc:ScientificName>
	<commonName xml:lang='pt'>Cação-anjo</commonName>
	<commonName xml:lang='en'>Angular angel shark</commonName>
	<commonName xml:lang='en'>Argentine angelshark</commonName>
	<commonName xml:lang='et'>Tähniline ingelhai</commonName>
	<commonName xml:lang='es'>Angelito</commonName>
	<commonName xml:lang='es'>Pez angel</commonName>
	<commonName xml:lang='es'>Pez ángel</commonName>
	<commonName xml:lang=''>Gebogen zee-engel</commonName>
	<commonName xml:lang=''>南美扁鯊</commonName>
	<commonName xml:lang=''>南美扁鲨</commonName>
	<synonym relationship='synonym'>Squatina punctata Marini, 1936</synonym>
	<dcterms:created>1999-09-13</dcterms:created>
	<dcterms:modified>2018-08-07</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=53932'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=53932'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=53932'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26375&amp;speccode=53932'>Carvalho-Filho, A.0 Peixes: costa Brasileira. Marca D'Agua, Sao Paolo, Brazil. 304 p. (Ref. 26375)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=53932'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=53932'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='2-8317-0564-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=36508&amp;speccode=53932'>Hilton-Taylor, C.0 2000 IUCN red list of threatened species. IUCN, Gland, Switzerland and Cambridge, UK. xviii + 61 p. (with 1 CD-ROM). (Ref. 36508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=53932'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=53932'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50838&amp;speccode=53932'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of January 2004. Database dump. Catalog databases as made available to FishBase in January 2004. (Ref. 50838)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52638&amp;speccode=53932'>Anonymous0 Fish collection of the Museu Oceanográfico do Vale do Itajaí, Universidade do Vale do Itajaí. Universidade do Vale do Itajaí. (Ref. 52638)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53443&amp;speccode=53932'>Soto, J.M.R.0 Annotated systematic checklist and bibliography of the coastal and oceanic fauna of Brazil. I. Sharks. Mare Magnum 1(1):51-119. (Ref. 53443)</reference>
	<reference isbn='9974-563-18-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=54736&amp;speccode=53932'>Nión, H., C. Ríos and P. Meneses0 Peces del Uruguay: Lista sistemática y nombres comunes. Montevideo, DINARA, Infopesca. (Ref. 54736)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55667&amp;speccode=53932'>Vögler, R., A.C. Milessi and R.A. Quiñones0 Trophic ecology of &lt;i&gt;Squatina guggenheim&lt;/i&gt; on the continental shelf off Uruguay and northern Argentina. J. Fish Biol. 62(6):1254-1267. (Ref. 55667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57911&amp;speccode=53932'>Bernardes, R.Á., J.L. de Figueiredo, A.R. Rodrigues, L.G. Fischer, C.M. Vooren, M. Haimovici and C.L.D.B. Rossi-Wongtschowski0 Peixes de zona econômica exclusiva da região sudeste-sul do Brasil: Levantamento com armadilhas, pargueiras e rede de arrasto de fundo. São Paulo: Editora da Universidade de São Paulo. 295 p. (Ref. 57911)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=53932'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58839&amp;speccode=53932'>Menni, R.C. and L.O. Lucifora0 Condrictios de la Argentina y Uruguay. &lt;i&gt;ProBiota&lt;/i&gt;, FCNyM, UNLP, Serie Técnica-Didáctica, La Plata, Argentina, 11:1-15. (Ref. 58839)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80860&amp;speccode=53932'>Menezes, N.A., P.A. Buckup, J.L. Figueiredo and R.L. Moura0 Catálogo das espécies de peixes marinhos do Brasil. São Paulo. Museu de Zoologia da Universidade de São Paulo, 160 pp. (Ref. 80860)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86510&amp;speccode=53932'>Colonello, J.H., L.O. Lucifora and A.M. Massa0 Reproduction of the angular angel shark (&lt;i&gt;Squatina guggenheim&lt;/i&gt;): geographic differences, reproductive cycle, and sexual dimorphism. ICES J. Mar. Sci. 64:131-140. (Ref. 86510)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87040&amp;speccode=53932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2011. (Ref. 87040)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=53932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95520&amp;speccode=53932'>Vaz, D.F.B. and M.R. De Carvalho0 Morphological and taxonomic revision of species of &lt;i&gt;Squatina&lt;/i&gt; from the Southwestern Atlantic Ocean (Chondrichthyes: Squatiniformes: Squatinidae). Zootaxa 3695(1):001-081. (Ref. 95520)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=53932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=53932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=53932'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=53932'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=53932'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='978-987-88-5833-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=130851&amp;speccode=53932'>Sánchez-Carnero, N., M.E. Góngora, M. Álvarez and A. Parma0 La pesca artesanal en Argentina: caminando las costas del país la. ed. 230. (Ref. 130851)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=131174&amp;speccode=53932'>Bunholi, I.V., B.L.D.S. Ferrette, R.R. Domingues, M.M. Rotundo, J.M. Cuevas, M. García, S. Gómez, R.H. Aché de Freitas, C. Oliveira, F. Foresti and F.F. Mendonça0 Multilocus phylogeography of the endemic and endangered angular angelshark (&lt;i&gt;Squatina guggenheim&lt;/i&gt;) in the southwest Atlantic Ocean. Hydrobiologia 849(9):2177-2192. (Ref. 131174)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132125&amp;speccode=53932'>Ferragut-Perello, F., S. Ramirez-Amaro, A.C. Tsikliras, N. Petit-Marty, D. Dimarchopoulou0 Exploitation and conservation status of the thornback ray (&lt;i&gt;Raja clavata&lt;/i&gt;) in the Balearic Islands (Western Mediterranean). Fishes 8(2):117. (Ref. 132125)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1999-09-15 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-10 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=53932</dc:source>
	<subject></subject>
	<dc:description>Bony fishes were the main food followed by crustaceans and molluscs (Ref. 55667). Ovoviviparous (Ref. 50449).  Adult males are characterized by the presence of a patch of small spines near the tip of the pectoral fins (Ref. 86510).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95520&amp;speccode=53932'>Vaz, D.F.B. and M.R. De Carvalho 2013 Morphological and taxonomic revision of species of &lt;i&gt;Squatina&lt;/i&gt; from the Southwestern Atlantic Ocean (Chondrichthyes: Squatiniformes: Squatinidae). Zootaxa 3695(1):001-081. (Ref. 95520)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2023-10-17 00:00:00</dcterms:created>
	<dcterms:modified>2023-10-17 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=53932</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>This species is distinguished by the following characters: with a single (or multiple in juveniles) row of enlarged, conical and morphologically distinct dermal denticles over dorsal longitudinal midline, from midlength of the pectoral fin base to origin of first dorsal fin (in some specimens, to origin of caudal fin); dorsal midline denticles are with enlarged and rounded basal plate, 3-4 times diameter of other dorsal denticles, crown with several median ridges (10-12), and crown apex acute and smooth, without ridges (vs. dorsal midline denticles on trunk morphologically similar to other trunk denticles and barely organized in a row in S. occulta and S. argentina); differs from S. occulta by having larger and fewer white blotches on dorsal surface, without surrounding blackish rings (i.e. absence of ocelli-like markings) and smaller in size at maturity, from 73.0-80.0 cm TL (vs. greater than 100.0 cm TL in S. occulta); differs from S. argentina by its ligther background colour, light to dark brown (vs. dark brown to reddish brown), with larger and fewer white blotches over dorsal surface, with a pair of enlarged, conical and morphologically distinct dermal denticles between spiracles (vs. interspiracular surface covered by smaller dorsal denticles, no enlarged denticles), the lower number of tooth rows with 18-22 vertical rows in both upper and lower jaws, tooth formula 9-9 to 11-11/9-9 to 11-11 (vs. 12-12/12-12 in i&gt;S. argentina), and with a straight external pectoral fin margin (vs. with convex anterior half of external margin).  Colouration: dorsal color pattern of light to dark brown background, with several white, creamy-white or yellowish large and rounded blotches variable in size (between one-half to 1.5 times eye-length) on entire dorsal surface (Ref. 95520).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95520&amp;speccode=53932'>Vaz, D.F.B. and M.R. De Carvalho 2013 Morphological and taxonomic revision of species of &lt;i&gt;Squatina&lt;/i&gt; from the Southwestern Atlantic Ocean (Chondrichthyes: Squatiniformes: Squatinidae). Zootaxa 3695(1):001-081. (Ref. 95520)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1999-09-15 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Southwest Atlantic:  southern Brazil and Argentina.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26375&amp;speccode=53932'>Carvalho-Filho, A. 1992 Peixes: costa Brasileira. Marca D'Agua, Sao Paolo, Brazil. 304 p. (Ref. 26375)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1999-09-15 00:00:00</dcterms:created>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>fisheries: commercial</dc:description>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=53932&amp;StockCode=44735</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Ovoviviparous, embryos feed solely on yolk (Ref. 50449).  Only the left ovary is functional (Ref. 57911).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=53932'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Grace Tolentino Pablico</agent>
	<dcterms:created>2005-10-06 00:00:00</dcterms:created>
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	<dc:source>guggenheim</dc:source>
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	<dc:description>Bony fishes were the main food followed by crustaceans and molluscs (Ref. 55667).  Diets of different size classes showed significant differences.  Male and female diets were not found to differ significantly (P&gt;0.01). Engraulis anchoita was the most consumed fish  (Ref. 55667).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55667&amp;speccode=53932'>Vögler, R., A.C. Milessi and R.A. Quiñones 2003 Trophic ecology of &lt;i&gt;Squatina guggenheim&lt;/i&gt; on the continental shelf off Uruguay and northern Argentina. J. Fish Biol. 62(6):1254-1267. (Ref. 55667)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
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	<dc:title>Max. size</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=53932</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>129 cm TL (male/unsexed; (Ref. 57911)); 130 cm TL (female)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57911&amp;speccode=53932'>Bernardes, R.Á., J.L. de Figueiredo, A.R. Rodrigues, L.G. Fischer, C.M. Vooren, M. Haimovici and C.L.D.B. Rossi-Wongtschowski 2005 Peixes de zona econômica exclusiva da região sudeste-sul do Brasil: Levantamento com armadilhas, pargueiras e rede de arrasto de fundo. São Paulo: Editora da Universidade de São Paulo. 295 p. (Ref. 57911)</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>demersal; brackish; marine; depth range 4 - 360 m (Ref. 57911), usually 4 - 266 m (Ref. 55667)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57911&amp;speccode=53932'>Bernardes, R.Á., J.L. de Figueiredo, A.R. Rodrigues, L.G. Fischer, C.M. Vooren, M. Haimovici and C.L.D.B. Rossi-Wongtschowski 2005 Peixes de zona econômica exclusiva da região sudeste-sul do Brasil: Levantamento com armadilhas, pargueiras e rede de arrasto de fundo. São Paulo: Editora da Universidade de São Paulo. 295 p. (Ref. 57911)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55667&amp;speccode=53932'>Vögler, R., A.C. Milessi and R.A. Quiñones 2003 Trophic ecology of &lt;i&gt;Squatina guggenheim&lt;/i&gt; on the continental shelf off Uruguay and northern Argentina. J. Fish Biol. 62(6):1254-1267. (Ref. 55667)</reference></dataObject>
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