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<taxon>
	<dc:identifier>FB-5446</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5446</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Acanthuriformes</dwc:Order>
	<dwc:Family>Chaetodontidae</dwc:Family>
	<dwc:Genus>Chaetodon</dwc:Genus>
	<dwc:ScientificName>Chaetodon kleinii Bloch, 1790</dwc:ScientificName>
	<commonName xml:lang='af'>Klein vlindervis</commonName>
	<commonName xml:lang=''>Alibangbang</commonName>
	<commonName xml:lang='da'>Kleins fanefisk</commonName>
	<commonName xml:lang='de'>Kleins Falterfisch</commonName>
	<commonName xml:lang='de'>Weißpunkt-Falterfisch</commonName>
	<commonName xml:lang='en'>Blacklip butterflyfish</commonName>
	<commonName xml:lang='en'>Bluehead butterflyfish</commonName>
	<commonName xml:lang='en'>Brown butterflyfish</commonName>
	<commonName xml:lang='en'>Butterflyfish</commonName>
	<commonName xml:lang='en'>Klein's butterfly</commonName>
	<commonName xml:lang='en'>Klein's butterfly fish</commonName>
	<commonName xml:lang='en'>Klein's butterflyfish</commonName>
	<commonName xml:lang='en'>Klein's coralfish</commonName>
	<commonName xml:lang='en'>Sunburst butterflyfish</commonName>
	<commonName xml:lang='en'>Whitespotted butterflyfish</commonName>
	<commonName xml:lang='en'>White-spotted butterflyfish</commonName>
	<commonName xml:lang='en'>Yellowspot butterflyfish</commonName>
	<commonName xml:lang='et'>Kleini harjashammas</commonName>
	<commonName xml:lang='fj'>daliga ragutu</commonName>
	<commonName xml:lang='fr'>Chétodon de klein</commonName>
	<commonName xml:lang='fr'>Chétodon de Klein</commonName>
	<commonName xml:lang='fr'>Papillon de Klein</commonName>
	<commonName xml:lang='fr'>Papillon de klein</commonName>
	<commonName xml:lang='fr'>Papillon jaune</commonName>
	<commonName xml:lang=''>Lauhau</commonName>
	<commonName xml:lang='ja'>Mizore chouchou-uo</commonName>
	<commonName xml:lang='ja'>Mizore-chô-chô-uo</commonName>
	<commonName xml:lang='ms'>Bagang</commonName>
	<commonName xml:lang='ms'>Bagang belang putih</commonName>
	<commonName xml:lang='ms'>Kepe orange</commonName>
	<commonName xml:lang='sm'>Tifitifi</commonName>
	<commonName xml:lang='tl'>Paru-paro</commonName>
	<commonName xml:lang='tl'>Paru-parung dagat</commonName>
	<commonName xml:lang='vi'>Cá Bướm nâu nhạt</commonName>
	<commonName xml:lang='es'>Mariposa de barra penumbre</commonName>
	<commonName xml:lang=''>Alanaasi bibee</commonName>
	<commonName xml:lang=''>Alibang-bang</commonName>
	<commonName xml:lang=''>Kalaykalay</commonName>
	<commonName xml:lang=''>Kapiged</commonName>
	<commonName xml:lang=''>Papillon</commonName>
	<commonName xml:lang=''>Pavillon</commonName>
	<commonName xml:lang=''>克氏蝴蝶魚</commonName>
	<commonName xml:lang=''>克氏蝴蝶鱼</commonName>
	<commonName xml:lang=''>珠蝴蝶魚</commonName>
	<commonName xml:lang=''>珠蝴蝶鱼</commonName>
	<commonName xml:lang=''>菠萝蝶</commonName>
	<commonName xml:lang=''>菠蘿蝶</commonName>
	<commonName xml:lang=''>蓝头蝶</commonName>
	<commonName xml:lang=''>麻包</commonName>
	<synonym relationship='synonym'>Anisochaetodon kleinii (Bloch, 1790)</synonym>
	<synonym relationship='synonym'>Chaetodon bellulus Thiollière, 1857</synonym>
	<synonym relationship='synonym'>Chaetodon cingulatus Fowler, 1934</synonym>
	<synonym relationship='synonym'>Chaetodon corallicola Snyder, 1904</synonym>
	<synonym relationship='synonym'>Chaetodon flavescens Bennett, 1831</synonym>
	<synonym relationship='synonym'>Chaetodon kleini Bloch, 1790</synonym>
	<synonym relationship='synonym'>Chaetodon melammystax Bloch &amp; Schneider, 1801</synonym>
	<synonym relationship='synonym'>Chaetodon melastomus Bloch &amp; Schneider, 1801</synonym>
	<synonym relationship='synonym'>Chaetodon virescens Cuvier, 1831</synonym>
	<synonym relationship='synonym'>Choetodon kleinii Bloch, 1790</synonym>
	<synonym relationship='synonym'>Exornator exornator kleinii (Bloch, 1790)</synonym>
	<synonym relationship='synonym'>Tetragonoptrus kleini (Bloch, 1790)</synonym>
	<dcterms:created>1992-05-07</dcterms:created>
	<dcterms:modified>2023-06-14</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=5446'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118793&amp;speccode=5446'>Allen, G.R. and M.V. Erdmann0 Coral reef fishes of Timor-Leste. p. 33-82.  In G.R. Allen and M.V. Erdmann (eds).  A rapid biological assessment of Timor-Leste. Conservation International. (Ref. 118793)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121724&amp;speccode=5446'>Songco, A.0 Personal communication. Tubbataha species list. Excel file "MyReport_Species_21May2019". (Ref. 121724)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121799&amp;speccode=5446'>University of Western Australia0 Pelagic BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 121799)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122752&amp;speccode=5446'>University of Western Australia0 Benthic  BRUVS data 2019. Video BRUVS from Jessica Meeuwig. (Ref. 122752)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124899&amp;speccode=5446'>van der Elst, Rudy, Simon Chater and Dennis King0 Fishes from Bassas Da India Atoll in the Mozambique Channel. Atoll Research Bulletin No. 629. (Ref. 124899)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126188&amp;speccode=5446'>Aranas, Kevin Nikko E.0 Reef Fishes Caught by Bobo Fish Traps in Barangay Daliao, Toril, Davao City Davao City (Philippines): University of Southern Philippines, B.Sc. Thesis, 161 p. (Ref. 126188)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=127348&amp;speccode=5446'>Grove, J.S., D.J. Long, D.R. Robertson and B.C. Victor0 List of fishes of the Galapagos Archipelago, Ecuador. J. Ocean Sci. Found. 39:14-22. (Ref. 127348)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=127519&amp;speccode=5446'>Hazes, B.0 Hazes' underwater photos. Collection of Philippine underwater photos. Unpublished. (Ref. 127519)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128523&amp;speccode=5446'>To, A.W.L., K.S.H. Ching and S.K.H. Shea0 Hong Kong reef fish photo guide. Eco-Education and Resources Centre (Ref. 128523)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128797&amp;speccode=5446'>Coleman, R., J.M. Copus, D.M. Coffey, R.K. Whitton and B.W. Bowen0 Shifting reef fish assemblages along a depth gradient in Pohnpei, Micronesia. PeerJ, 6: e4650. (Ref. 128797)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=5446'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130287&amp;speccode=5446'>Balisco, R.A.T., V.S. Ticzon, B.R. Samaniego, W.-C. Huang, B.J. Gonzales and T.-Y. Liao0 Marine fishes of Palawan, Philippines: species diversity, new records, and conservation status. Regional Studies in Marine Science 60(2023) 102825:1-10. (Ref. 130287)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=5446'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138103&amp;speccode=5446'>Petit-Marty, N., M. Vázquez-Luis and I.E. Hendriks0 Use of the nucleotide diversity in COI mitochondrial gene as an early diagnostic of conservation status of animal species. Conserv. Lett. 14(1):1-7. (Ref. 138103)</reference><dataObject>
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	<dc:description>Adults occur in deeper lagoons and channels, and seaward reefs (Ref. 1602).  Benthopelagic (Ref. 58302).  Depth 2-61 m, usually below 10 m (Ref. 90102).  They Occur singly or in pairs (Ref. 37816).  Common, omnivorous individuals that feed mainly on soft coral polyps (mainly on Sarcophyton tracheliophorum and Litophyton viridis), algae and zooplankton.  Oviparous (Ref. 205).  Form pairs during breeding (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1602&amp;speccode=5446'>Myers, R.F. 1991 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602)</reference></dataObject><dataObject>
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	<dc:description>Body is yellowish brown with two broad white vertical bars running across the body one from near the origin of the dorsal spine and the other from the middle of the back.  A black bar runs vertically across the eye.  There are numerous dotted horizontal stripes on the sides.  The margin of caudal fin is transparent (Ref. 4855).  Snout length 2.5-3.2 in HL.  Body depth 1.5-1.8 in SL (Ref. 90102).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4855'>Burgess, W.E. 1978 Butterflyfishes of the world. A monograph of the Family Chaetodontidae. T.F.H. Publications, Neptune City, New Jersey.</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 13 - 14; Dorsal soft rays (total): 20 - 23;  Analspines: 3; Analsoft rays: 17 - 20; Vertebrae: 24</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4855'>Burgess, W.E. 1978 Butterflyfishes of the world. A monograph of the Family Chaetodontidae. T.F.H. Publications, Neptune City, New Jersey.</reference></dataObject><dataObject>
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	<dc:description>Indo-West Pacific:  Red Sea and East Africa (south to Coffee Bay, South Africa, Ref. 5372) to the Hawaiian Islands and Samoa, north to southern Japan, south to New South Wales, Australia and New Caledonia.  Eastern Pacific:  Galapagos Islands (Ref. 5227).</dc:description>
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	<reference isbn='0-8248-1346-4.' url='http://www.fishbase.org/references/FBRefSummary.php?id=2334&amp;speccode=5446'>Randall, J.E., G.R. Allen and R.C. Steene 1990 Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334)</reference></dataObject><dataObject>
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	<dc:description>fisheries: subsistence fisheries; aquarium: commercial</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4907&amp;speccode=5446'>Edwards, A.J. and A.D. Shepherd 1992 Environmental implications of aquarium-fish collection in the Maldives, with proposals for regulation. Environ. Conserv. 19:61-72. (Ref. 4907)</reference></dataObject><dataObject>
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	<dc:description>Distinct pairing (Ref. 205). Monogamous mating is observed as both obligate and social (Ref. 52884).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=240&amp;speccode=5446'>Thresher, R.E. 1984 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240)</reference></dataObject><dataObject>
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	<dc:source>kleinii</dc:source>
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	<dc:description>Occur in deeper lagoons and channels, and seaward reefs (Ref. 1602).  Occur singly or in pairs (Ref. 37816).  Common, omnivorous individuals that feed mainly on soft coral polyps (mainly on Sarcophyton tracheliophorum and Litophyton viridis), algae and zooplankton.  Feed on plants, benthic and planktonic invertebrates (Ref. 6110, 13550).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4859&amp;speccode=5446'>Steene, R.C. 1978 Butterfly and angelfishes of the world. A.H. &amp; A.W. Reed Pty Ltd., Australia. vol. 1. 144 p. (Ref. 4859)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>15.0 cm TL (male/unsexed; (Ref. 5372))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5372&amp;speccode=5446'>Heemstra, P.C. 1986 Chaetodontidae. p. 627-632. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 5372)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5446</dc:identifier>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1992-05-07 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-20 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5446</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>reef-associated; non-migratory (Ref. 52884); marine; depth range 0 - 90 m (Ref. 128797), usually 10 - ? m (Ref. 1602)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52884&amp;speccode=5446'>Whiteman, E.A. and I.M. Côté 2004 Monogamy in marine fishes. Biol. Rev. 79:351-375. (Ref. 52884)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128797&amp;speccode=5446'>Coleman, R., J.M. Copus, D.M. Coffey, R.K. Whitton and B.W. Bowen 2018 Shifting reef fish assemblages along a depth gradient in Pohnpei, Micronesia. PeerJ, 6: e4650. (Ref. 128797)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1602&amp;speccode=5446'>Myers, R.F. 1991 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602)</reference></dataObject>
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