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	<dc:identifier>FB-5460</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Gobiiformes</dwc:Order>
	<dwc:Family>Gobiidae</dwc:Family>
	<dwc:Genus>Pandaka</dwc:Genus>
	<dwc:ScientificName>Pandaka pygmaea Herre, 1927</dwc:ScientificName>
	<commonName xml:lang='da'>Pygmækutling</commonName>
	<commonName xml:lang='de'>Zwerggrundel</commonName>
	<commonName xml:lang='en'>Dwarf pygmy goby</commonName>
	<commonName xml:lang='en'>Dwarf Pygmy Goby</commonName>
	<commonName xml:lang='et'>Harilik pandaka</commonName>
	<commonName xml:lang='fi'>Kääpiötokko</commonName>
	<commonName xml:lang='sv'>Pandaka</commonName>
	<commonName xml:lang='tl'>Bia</commonName>
	<commonName xml:lang=''>Pritlikavi glavač</commonName>
	<commonName xml:lang=''>矮鰕虎魚</commonName>
	<commonName xml:lang=''>矮鰕虎鱼</commonName>
	<dcterms:created>1992-05-13</dcterms:created>
	<dcterms:modified>2024-01-25</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5460'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=280&amp;speccode=5460'>Herre, A.W.C.T.0 Check list of Philippine fishes. Res. Rep. U.S. Fish Wild. Serv. (20):1-977. (Ref. 280)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=419&amp;speccode=5460'>Koumans, F.P.0 Gobioidea. In M. Weber and L. de Beaufort (eds.) Fishes of the Indo-Australian Archipelago, Vol. 10. E.J. Brill, Leiden. 423 p. (Ref. 419)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=439&amp;speccode=5460'>Herre, A.W.C.T.0 The Philippine Bureau of Science monographic publications on fishes: gobies of the Philippines and China Sea. Monogr. Bur. Sci. Manila 23:1-352. (Ref. 439)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2060&amp;speccode=5460'>Baensch, H.A. and R. Riehl0 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2857&amp;speccode=5460'>Herre, A.W.C.T. and A.F. Umali0 English and local common names of Philippine fishes. U. S. Dept. of Interior and Fish and Wildl. Serv. Circular No. 14, U. S. Gov't Printing Office, Washington. 128 p. (Ref. 2857)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4924&amp;speccode=5460'>Sarenas, A.M. and I.A. Ronquillo0 Discovery of living specimens and re-description of &lt;i&gt;Pandaka pygmea&lt;/i&gt; Herre. Philipp. J. Sci. 83(3):257-261. (Ref. 4924)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6216&amp;speccode=5460'>Norman, A.0 The smallest fish there is. Freshwat. Mar. Aquar. 4(1):26-27. (Ref. 6216)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6218&amp;speccode=5460'>Liu, R.K. and R.L. Walford0 Observations on the lifespans of several species of annual fishes and of the world's smallest fishes. Exp. Geront. 5:241-246. (Ref. 6218)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6376&amp;speccode=5460'>IUCN0 1994 IUCN red list of threatened animals. International Union for Conservation of Nature and Natural Resources, Gland, Switzerland and Cambridge, U.K. (Ref. 6376)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7050&amp;speccode=5460'>Kottelat, M., A.J. Whitten, S.N. Kartikasari and S. Wirjoatmodjo0 Freshwater fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong. 221 p. (Ref. 7050)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7463&amp;speccode=5460'>Nelson, J.S.0 Fishes of the world. Third edition. John Wiley &amp; Sons, Inc., New York. 600 p. (Ref. 7463)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12165&amp;speccode=5460'>Bleher, H.0 Bombon. Aqua geographia 12(4):6-34. (Ref. 12165)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12255&amp;speccode=5460'>Baillie, J. and B. Groombridge (eds.)0 1996 IUCN red list of threatened animals. IUCN, Gland, Switzerland. 378 p. (Ref. 12255)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=5460'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=5460'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=5460'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='2-8317-0564-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=36508&amp;speccode=5460'>Hilton-Taylor, C.0 2000 IUCN red list of threatened species. IUCN, Gland, Switzerland and Cambridge, UK. xviii + 61 p. (with 1 CD-ROM). (Ref. 36508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36648&amp;speccode=5460'>Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=5460'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=5460'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=5460'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43309&amp;speccode=5460'>Anonymous0 Fish collection of the University of the Philippines in the Visayas Museum. UPV Museum. (Ref. 43309)</reference>
	<reference isbn='0-7307-1292-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43716&amp;speccode=5460'>Larson, H.K.0 A revision of the gobiid fish genus &lt;i&gt;Mugilogobius&lt;/i&gt; (Teleostei: Gobioidei), and its systematic placement. Rec. West. Aust. Mus. (Suppl. No. 62):1-233. (Ref. 43716)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46888&amp;speccode=5460'>McDowall, R.M.0 The evolution of diadromy in fishes (revisited) and its place in phylogenetic analysis. Rev. Fish Biol. Fish. 7(4):443-462. (Ref. 46888)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47438&amp;speccode=5460'>Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438)</reference>
	<reference isbn='1-881173-36-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=48613&amp;speccode=5460'>Werner, T.B. and G.R. Allen0 A rapid marine biodiversity assessment of the Calamianes Islands, Palawan province, Philippines. RAP Bulletin of Biological Assessment 17. Washington, D.C.:Conservation International. (Ref. 48613)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=5460'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=5460'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=5460'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=5460'>Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference>
	<reference isbn='9781578084364' url='http://www.fishbase.org/references/FBRefSummary.php?id=92840&amp;speccode=5460'>Patzner, R.A., J.L. Van Tassell, M. Kovačić and B.G. Kapoor0 The biology of gobies. Enfield, NH : Science Publishers ; Boca Raton, FL : DIstributed by CRC Press, 685 p. (Ref. 92840)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94476&amp;speccode=5460'>Kottelat, M.0 The fishes of the inland waters of Southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries. The Raffles Bulletin of Zoology 2013 (Suppl. 27):1-663. (Ref. 94476)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=5460'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='2350-4757' url='http://www.fishbase.org/references/FBRefSummary.php?id=128520&amp;speccode=5460'>Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=7' logoURL='' role='photographer'>Robbie N. Cada</agent>
	<dcterms:created>1993-07-01</dcterms:created>
	<dcterms:modified>2008-06-25</dcterms:modified>
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	<mediaURL>http://www.fishbase.us/images/species/Papyg_u0.gif</mediaURL>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1992-05-13 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dc:description>This is one of the smallest freshwater fishes:  males are mature at 9 mm, females at 15 mm.  The species used to frequent shady river banks in the Malabon River, Rizal Province, Luzon, Philippines; however, the type locality has been reclaimed and the remaining waters in the area are heavily polluted; the species is considered extinct in the Philippines (Rainer Froese, pers. comm., 2004).  It has been apparently imported into Germany in 1958;  a color photo of the fish was taken in an aquarium (Ref. 2060).  Found in brackish waters and mangrove areas of Indonesia (M. Kottelat, pers. comm.).</dc:description>
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	<reference isbn='0-7307-1292-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=43716&amp;speccode=5460'>Larson, H.K. 2001 A revision of the gobiid fish genus &lt;i&gt;Mugilogobius&lt;/i&gt; (Teleostei: Gobioidei), and its systematic placement. Rec. West. Aust. Mus. (Suppl. No. 62):1-233. (Ref. 43716)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1992-09-25 00:00:00</dcterms:created>
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	<dc:description>Dark spots on sides of body somewhat forming 4 cross-bands; heavy pigmentations on all bases of fins, except ventrals, apparently are continuation of body pigmentation.  Head and nape naked.  22 to 25 scales in longitudinal series (Ref. 4924); further characterized by presence of black spot at middle of caudal fin base; first dorsal fin yellowish with black anterior; rounded caudal fin (Ref. 90102).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4924&amp;speccode=5460'>Sarenas, A.M. and I.A. Ronquillo 1955 Discovery of living specimens and re-description of &lt;i&gt;Pandaka pygmea&lt;/i&gt; Herre. Philipp. J. Sci. 83(3):257-261. (Ref. 4924)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
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	<dc:description>Dorsal spines (total): 7; Dorsal soft rays (total): 6 - 7;  Analspines: 1; Analsoft rays: 6 - 7</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4924&amp;speccode=5460'>Sarenas, A.M. and I.A. Ronquillo 1955 Discovery of living specimens and re-description of &lt;i&gt;Pandaka pygmea&lt;/i&gt; Herre. Philipp. J. Sci. 83(3):257-261. (Ref. 4924)</reference></dataObject><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1992-05-13 00:00:00</dcterms:created>
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	<dc:description>Asia:  Indonesia and Philippines.  Originally reported from the Malabon River, Rizal, Philippines, which has meanwhile been reclaimed.  According to Ref. 6216 it has also been collected in the sea at Culion Island, off Palawan, Philippines.  Recently collected in Bali (1991), Sulawesi (1988), and Singapore (1992), M. Kottelat, pers. comm.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=280&amp;speccode=5460'>Herre, A.W.C.T. 1953 Check list of Philippine fishes. Res. Rep. U.S. Fish Wild. Serv. (20):1-977. (Ref. 280)</reference></dataObject><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2060&amp;speccode=5460'>Baensch, H.A. and R. Riehl 1985 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060)</reference></dataObject><dataObject>
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	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
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	<dc:description>Amphidromous. Refers to fishes that regularly migrate between freshwater and the sea (in both directions), but not for the purpose of breeding, as in anadromous and catadromous species. Sub-division of diadromous.  Migrations should be cyclical and predictable and cover more than 100 km.Characteristic elements in amphidromy are: reproduction in fresh water, passage to sea by newly hatched larvae, a period of feeding and growing at sea usually a few months long, return to fresh water of well-grown juveniles, a further period of feeding and growing in fresh water, followed by reproduction there (Ref. 82692).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46888&amp;speccode=5460'>McDowall, R.M. 1997 The evolution of diadromy in fishes (revisited) and its place in phylogenetic analysis. Rev. Fish Biol. Fish. 7(4):443-462. (Ref. 46888)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
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	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=5460&amp;StockCode=5731</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Probably a non-guarder (RF).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5460'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-5460</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1992-05-13 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5460</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>1.1 cm SL (male/unsexed; (Ref. 2060)); 1.5 cm TL (female)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2060&amp;speccode=5460'>Baensch, H.A. and R. Riehl 1985 Aquarien atlas. Band 2. Mergus, Verlag für Natur-und Heimtierkunde GmbH, Melle, Germany. 1216 p. (Ref. 2060)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5460</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='http://www.fishbase.de/rfroese' logoURL='' role='recorder'>Rainer Froese</agent>
	<dcterms:created>1992-05-13 00:00:00</dcterms:created>
	<dcterms:modified>2024-01-25 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5460</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>demersal; amphidromous (Ref. 46888); freshwater; brackish; marine; pH range: 7.0 - 8.4; dH range: 30; depth range 0 - 2 m (Ref. 90102)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46888&amp;speccode=5460'>McDowall, R.M. 1997 The evolution of diadromy in fishes (revisited) and its place in phylogenetic analysis. Rev. Fish Biol. Fish. 7(4):443-462. (Ref. 46888)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=5460'>Allen, G.R. and M.V. Erdmann 2012 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference></dataObject>
</taxon>
</response>