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<taxon>
	<dc:identifier>FB-5509</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5509</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rhinopristiformes</dwc:Order>
	<dwc:Family>Rhinobatidae</dwc:Family>
	<dwc:Genus>Acroteriobatus</dwc:Genus>
	<dwc:ScientificName>Acroteriobatus annulatus (Smith, 1841)</dwc:ScientificName>
	<commonName xml:lang='pl'>Roszak</commonName>
	<commonName xml:lang='af'>Banjo</commonName>
	<commonName xml:lang='af'>Kleiner sandkruiper</commonName>
	<commonName xml:lang='af'>Sandhaai</commonName>
	<commonName xml:lang='ar'>Barbar</commonName>
	<commonName xml:lang='ar'>Fanto</commonName>
	<commonName xml:lang='cs'>Pilohrbet kroužkový</commonName>
	<commonName xml:lang='cs'>Rejnokovec kruhový</commonName>
	<commonName xml:lang='en'>Guitarfish</commonName>
	<commonName xml:lang='en'>Lesser guitarfish</commonName>
	<commonName xml:lang='en'>Lesser sandshark</commonName>
	<commonName xml:lang='en'>Sandshark</commonName>
	<commonName xml:lang='et'>Mummuline logardrai</commonName>
	<commonName xml:lang='fr'>Poisson-guitare du Cap</commonName>
	<commonName xml:lang='es'>Guitarra del Cabo</commonName>
	<commonName xml:lang=''>Kleine gitaarrog</commonName>
	<commonName xml:lang=''>突吻犁头鳐</commonName>
	<commonName xml:lang=''>突吻犁頭鰩</commonName>
	<synonym relationship='synonym'>Rhinobatos annulatus Smith, 1841</synonym>
	<synonym relationship='misapplied name'>Rhinobatos columnae (non Bonaparte, 1836)</synonym>
	<synonym relationship='synonym'>Rhinobatus annulatus Smith, 1841</synonym>
	<dcterms:created>1992-06-02</dcterms:created>
	<dcterms:modified>2022-01-22</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5509'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3670&amp;speccode=5509'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. C. Struik, Cape Town. 367 p. (Ref. 3670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3678&amp;speccode=5509'>Torres, F.S.B. Jr.0 Tabular data on marine fishes from Southern Africa, Part I. Length-weight relationships. Fishbyte 9(1):50-53. (Ref. 3678)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5509'>Compagno, L.J.V.0 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4332&amp;speccode=5509'>van der Elst, R.P. and F. Adkin (eds.)0 Marine linefish: priority species and research objectives in southern Africa. Oceanogr. Res. Inst., Spec. Publ. No.1. 132 p. (Ref. 4332)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=5509'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5509'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10311&amp;speccode=5509'>Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman &amp; Hall, London. (Ref. 10311)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12484&amp;speccode=5509'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. (Ref. 12484)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=5509'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27121&amp;speccode=5509'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265p. (Ref. 27121)</reference>
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	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=5509'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
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	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=5509'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
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	<reference isbn='92-5-104345-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=36731&amp;speccode=5509'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265 p. (Ref. 36731)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36992&amp;speccode=5509'>Whitfield, A.K. and S.J.M. Blaber0 Predation on striped mullet (&lt;i&gt;Mugil cephalus&lt;/i&gt;) by &lt;i&gt;Crocodylus niloticus&lt;/i&gt; at St. Lucia, South Africa. Copeia 1979(2):266-269. (Ref. 36992)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39358&amp;speccode=5509'>Harris, S.A., B.A. Bennett and G.M. Branch0 An assessment of the role of the sand shark &lt;i&gt;Rhinobatos annulatus&lt;/i&gt; as a predator in Langebaan Lagoon. S. Afr. J. mar. Sci. 7:153-159. (Ref. 39358)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=5509'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=5509'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47487&amp;speccode=5509'>Bundesanstalt für Landwirtschaft und Ernährung0 Verzeichnis der Handelsbezeichnungen für Erzeugnisse der Fischerei und der Aquakultur. Retrieved 17 June 2003, from http://www.ble.de. (Ref. 47487)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=5509'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51054&amp;speccode=5509'>Hanel, L. and J. Novák0 České názvy zivočichů V. Ryby a rybovití obratlovci (Pisces) II., nozdratí (Sarcopterygii), paprskoploutví (Actinopterygii) [chrupavčití (Chondrostei), kostnatí (Neopterygii): kostlíni (Semionotiformes) - bezostní (Clupeiformes)]. Národní muzeum (zoologické odděleí), Praha. (Ref. 51054)</reference>
	<reference isbn='80-7036-139-5' url='http://www.fishbase.org/references/FBRefSummary.php?id=51055&amp;speccode=5509'>Hanel, L. and J. Novák0 České názvy zivočichů V. Ryby a ryboviti obratlovci (Pisces) 3., maloústí (Gonorhynchiformes) - máloostní (Cypriniformes). Národní muzeum (zoologické oddělení), Praha. (Ref. 51055)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=5509'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=5509'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=5509'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=5509'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=5509'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=5509'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=5509'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118717&amp;speccode=5509'>Tsuboi, M., W. van der Bijl, B.T. Kopperud, J. Erritzoe, K.L. Voje, A. Kotrschal, K.E. Yopak, S.P. Collin, A.N. Iwaniuk and N. Kolm0 Breakdown of brain-body allometry and the encephalization of birds and mammals. Nat. Ecol. Evol. 2:1495-1500. (Ref. 118717)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123477&amp;speccode=5509'>ECoF, Eschmeyer's Catalog of Fishes0 Catalog of Fishes. 1985 - onwards. &lt;b&gt;CITATION NOT TO BE USED OUTSIDE FISHBASE.  SEE REMARKS&lt;/b&gt;. Authorship varies in time: Eschmeyer, W.N.; Fricke, R.; Van der Laan, R.  Web database. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (Ref. 123477)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124054&amp;speccode=5509'>Vaughan. D.B. and L.A. Chisholm0 A new species of &lt;i&gt;Neoheterocotyle&lt;/i&gt; Hargis, 1955 (Monogenea: Monocotylidae) from the gills of &lt;i&gt;Rhinobatos annulatus&lt;/i&gt; Müller &amp; Henle (Rhinobatidae) off the southern tip of Africa. Syst. Parasitol. 77:205-213. (Ref. 124054)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124055&amp;speccode=5509'>Vaughan, D.B. and L.A. Chisholm0 Amendment of &lt;i&gt;Pseudoleptobothrium&lt;/i&gt; Young, 1967 (Monogenea, Microbothriidae) with the description of &lt;i&gt;Pseudoleptobothrium christisoni&lt;/i&gt; sp. nov. from the dermal denticles of &lt;i&gt;Rhinobatos annulatus&lt;/i&gt; (Rhinobatidae) off the southern tip of Africa. Acta Parasitologica 56(3):280-289. (Ref. 124055)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5509'>Weigmann, S., D.A. Ebert, B. Séret0 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference>
	<reference isbn='978-1-990951-28-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=127960&amp;speccode=5509'>Heemstra, P.C., E. Heemstra, D.A. Ebert, W. Holleman and J.E. Randall (eds.)0 Coastal fishes of the Western Indian Ocean Vol. 1. South African Institute for Aquatic Biodiversity, a National Research Facility of the National Research Foundation (NRF-SAIAB), 730 p. (Ref. 127960)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128078&amp;speccode=5509'>CITES0 Appendices I, II and III valid from 11 January 2023 and 25 November 2023. UNEP. (Ref. 128078)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=131153&amp;speccode=5509'>CITES0 Appendices I, II and III. UNEP. (Ref. 131153)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138356&amp;speccode=5509'>Groeneveld, M.J., J.D. Klein, M. van Staden, R.H. Bennett, M.L. Dicken, D.A. Ebert, B.Q. Mann, K. Perumal, R.G.A. Watson and A.E. Bester-van der Merwe0 Molecular taxonomy of guitarfishes (Rhinobatidae: Acroteriobatus). Research Square online. 2025. (Ref. 138356)</reference><dataObject>
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	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
	<dcterms:modified>2022-01-22 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:description>Found from the surf line to about 70 m depth (Ref. 5578), also reported as common in shallow waters to at least 75 m depth (Ref. 114953); also enters estuaries (Ref. 27121).  Feeds on crustaceans, bivalves, polychaete worms and small bony fishes (Ref. 5578).  Ovoviviparous (Ref. 50449). Males mature at ca. 59 cm TL, females at 62-65 cm TL; born at ca. 23 cm TL. Produces litters of 2-10 pups (Ref. 114953). Taken by surf anglers (Ref. 3919).  Flesh highly esteemed (Ref. 3919). It is parasitised by the monogeneans Pseudoleptobothrium christisoni on the dorsal skin surface (Ref. 124055) and Neoheterocotyle robii on the gill lamellae (Ref. 124054).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5509'>Compagno, L.J.V. 1986 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference></dataObject><dataObject>
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	<dc:description>Southeast Atlantic to West Indian Ocean: Angola to South Africa and southern Mozambique.</dc:description>
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	<reference isbn='978-1-990951-28-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=127960&amp;speccode=5509'>Heemstra, P.C., E. Heemstra, D.A. Ebert, W. Holleman and J.E. Randall (eds.) 2022 Coastal fishes of the Western Indian Ocean Vol. 1. South African Institute for Aquatic Biodiversity, a National Research Facility of the National Research Foundation (NRF-SAIAB), 730 p. (Ref. 127960)</reference></dataObject><dataObject>
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	<dc:description>fisheries: commercial; gamefish: yes; aquarium: commercial</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5509'>Compagno, L.J.V. 1986 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5509'>Compagno, L.J.V. 1986 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124055&amp;speccode=5509'>Vaughan, D.B. and L.A. Chisholm 2011 Amendment of &lt;i&gt;Pseudoleptobothrium&lt;/i&gt; Young, 1967 (Monogenea, Microbothriidae) with the description of &lt;i&gt;Pseudoleptobothrium christisoni&lt;/i&gt; sp. nov. from the dermal denticles of &lt;i&gt;Rhinobatos annulatus&lt;/i&gt; (Rhinobatidae) off the southern tip of Africa. Acta Parasitologica 56(3):280-289. (Ref. 124055)</reference></dataObject><dataObject>
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	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449).  2-10 young born in the summer (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5509'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<dc:source>annulatus</dc:source>
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	<dc:description>Inhabits shallow water near the surf zone.  Feeds on benthic organisms (Ref. 9773).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4332&amp;speccode=5509'>van der Elst, R.P. and F. Adkin (eds.) 1991 Marine linefish: priority species and research objectives in southern Africa. Oceanogr. Res. Inst., Spec. Publ. No.1. 132 p. (Ref. 4332)</reference></dataObject><dataObject>
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	<dc:description>140 cm TL (male/unsexed; (Ref. 3919))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5509'>Compagno, L.J.V. 1986 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5509</dc:source>
	<dc:description>demersal; brackish; marine; depth range 75 - ? m (Ref. 114953)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=5509'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject>
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