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	<dc:identifier>FB-5510</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5510</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rhinopristiformes</dwc:Order>
	<dwc:Family>Rhinobatidae</dwc:Family>
	<dwc:Genus>Acroteriobatus</dwc:Genus>
	<dwc:ScientificName>Acroteriobatus leucospilus (Norman, 1926)</dwc:ScientificName>
	<commonName xml:lang='af'>Gryskol-sandkruiper</commonName>
	<commonName xml:lang='da'>Gråplettet guitarfisk</commonName>
	<commonName xml:lang='en'>Blue-spotted guitarfish</commonName>
	<commonName xml:lang='en'>Grayspot guitarfish</commonName>
	<commonName xml:lang='en'>Grayspottted guitarfish</commonName>
	<commonName xml:lang='en'>Greyspot guitarfish</commonName>
	<commonName xml:lang='en'>Greyspot sandshark</commonName>
	<commonName xml:lang='et'>Heletäpiline logardrai</commonName>
	<commonName xml:lang='fr'>Raie guitare à taches bleues</commonName>
	<commonName xml:lang='mg'>Soroboa</commonName>
	<commonName xml:lang=''>Grijsgestippelde gitaarrog</commonName>
	<commonName xml:lang=''>灰斑犁头鳐</commonName>
	<commonName xml:lang=''>灰斑犁頭鰩</commonName>
	<synonym relationship='misapplied name'>Rhinobatos blochi (non Müller &amp; Henle, 1841)</synonym>
	<synonym relationship='synonym'>Rhinobatos leucospilus Norman, 1926</synonym>
	<synonym relationship='synonym'>Rhinobatus leucospilus Norman, 1926</synonym>
	<dcterms:created>1992-06-02</dcterms:created>
	<dcterms:modified>2021-07-08</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5510'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3670&amp;speccode=5510'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. C. Struik, Cape Town. 367 p. (Ref. 3670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3678&amp;speccode=5510'>Torres, F.S.B. Jr.0 Tabular data on marine fishes from Southern Africa, Part I. Length-weight relationships. Fishbyte 9(1):50-53. (Ref. 3678)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3919&amp;speccode=5510'>Compagno, L.J.V.0 Rhinobatidae. p. 128-131. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 3919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=5510'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='0-913235-53-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=4537&amp;speccode=5510'>Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5510'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10311&amp;speccode=5510'>Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman &amp; Hall, London. (Ref. 10311)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11228&amp;speccode=5510'>Heemstra, P.C.0 Additions and corrections for the 1995 impression. p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin. (Ref. 11228)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12484&amp;speccode=5510'>van der Elst, R.0 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. (Ref. 12484)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=5510'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=5510'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=5510'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=5510'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=5510'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51471&amp;speccode=5510'>Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=5510'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='978-2-7592-0045-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=81519&amp;speccode=5510'>Taquet, M. and A. Diringer0 Poissons de l'Océan Indien et de la mer Rouge. Versailles (France): Éditions Quæ. 527 p. (Ref. 81519)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=5510'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=5510'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=5510'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret0 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=128078&amp;speccode=5510'>CITES0 Appendices I, II and III valid from 11 January 2023 and 25 November 2023. UNEP. (Ref. 128078)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=131153&amp;speccode=5510'>CITES0 Appendices I, II and III. UNEP. (Ref. 131153)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=138356&amp;speccode=5510'>Groeneveld, M.J., J.D. Klein, M. van Staden, R.H. Bennett, M.L. Dicken, D.A. Ebert, B.Q. Mann, K. Perumal, R.G.A. Watson and A.E. Bester-van der Merwe0 Molecular taxonomy of guitarfishes (Rhinobatidae: Acroteriobatus). Research Square online. 2025. (Ref. 138356)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
	<dcterms:modified>2021-07-08 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5510</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Found inshore from the surf line to a depth of 100 m (Ref. 5578, 11228).  Feeds on bivalves, gastropods, shrimps, crabs and small bony fishes (Ref. 5578).  Common length 92-96 cm TL; males and females mature at about 56 cm TL and birth size ca. 25 cm TL (Ref. 114953).   Ovoviviparous (Ref. 50449).  Flesh esteemed (Ref. 3919).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2021-07-08 00:00:00</dcterms:created>
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	<dc:description>This medium-sized species is distinguished by the following set of characters: dorsal surface smooth, no prominent thorns or tubercles, except for slightly enlarged granular denticles partially around orbital rims, and rather regularly distributed along midline from nape to near first dorsal-fin origin, none between dorsal fins and upper caudal fin; snout semi-translucent with elongated bluish spots giving stripe-nosed appearance, numerous small bluish spots covering the snout, pectoral, pelvic, dorsal, and caudal fins but not the central disc, on a sandy brown background with darker brown spots of varying sizes covering the disc, tail, fin bases, and fins, and sometimes giving the caudal peduncle a striped appearance, the outer edges of pectoral- and pelvic-fin margins blue, while lateral tail folds white or striped blue and brown and ventral surface white; nasal lamellae 37-41; upper jaw tooth row count  ca. 60-75; post-synarcual centra 187-192; total vertebral segments 200-205; total pectoral skeleton radials 63-70 (Ref. 124452).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2021-07-08 00:00:00</dcterms:created>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Vertebrae: 200 - 205</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-06-02 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Western Indian Ocean: South Africa, Mozambique, and Tanzania.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
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	<dc:title>Importance</dc:title>
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	<dc:description>fisheries: subsistence fisheries; gamefish: yes</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12484&amp;speccode=5510'>van der Elst, R. 1993 A guide to the common sea fishes of southern Africa. (3rd Ed.). Struik Publishers, Cape Town. 398 p. (Ref. 12484)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=5510'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=5510&amp;StockCode=5799</dc:source>
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	<dc:description>Exhibit ovoviparity (aplacental viviparity), with embryos feeding initially on yolk, then receiving additional nourishment from the mother by indirect absorption of uterine fluid enriched with mucus, fat or protein through specialised structures (Ref. 50449).  Bears 2 to 4 young (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5510'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5510</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>56.5 cm TL (male/unsexed; (Ref. 124452)); 96 cm TL (female)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>demersal; marine; depth range 0 - 100 m (Ref. 124452), usually ? - 37 m (Ref. 124452)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=124452&amp;speccode=5510'>Weigmann, S., D.A. Ebert, B. Séret 2021 Resolution of the &lt;i&gt;Acroteriobatus leucospilus&lt;/i&gt; species complex, with a redescription of &lt;i&gt;A. leucospilus&lt;/i&gt; (Norman, 1926) and descriptions of two new western Indian Ocean species of &lt;i&gt;Acroteriobatus&lt;/i&gt; (Rhinopristiformes, Rhinobatidae). Marine Biodiversity (2021):51:58. (Ref. 124452)</reference></dataObject>
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