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	<dc:identifier>FB-55511</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55511</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Gobiiformes</dwc:Order>
	<dwc:Family>Oxudercidae</dwc:Family>
	<dwc:Genus>Apocryptodon</dwc:Genus>
	<dwc:ScientificName>Apocryptodon punctatus Tomiyama, 1934</dwc:ScientificName>
	<commonName xml:lang='en'>Small-spine tadpole-goby</commonName>
	<commonName xml:lang='ru'>пуголовка Махмудбеева</commonName>
	<dcterms:created>2000-03-22</dcterms:created>
	<dcterms:modified>2014-02-27</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26334&amp;speccode=55531'>Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov0 An annotated check-list of the freshwater fishes of Russia. J. Ichthyol. 37(9):687-736. (Ref. 26334)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27116&amp;speccode=55531'>McAllister, D.E.0 A working list of fishes of the world. Copies available from D.E. McAllister, Canadian Museum of Nature, P.O. Box 3443, Ottawa, Ontario K1P 6P4, Canada. 2661 p. plus 1270 p. Index. (Ref. 27116)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33021&amp;speccode=55531'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of November 1999. Catalog databases as made available to FishBase in November 1999. (Ref. 33021)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=45043&amp;speccode=55531'>Bogutskaya, N.G.0 Common names of Russian freshwater fishes. Spreadsheet with cyrillic names made available for inclusion in FishBase. Unpublished. (Ref. 45043)</reference>
	<reference isbn='5-02-002872-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=50519&amp;speccode=55531'>Reshetnikov, Y.S.0 Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58297&amp;speccode=55531'>Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Volga River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58297)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=55531'>Kottelat, M. and J. Freyhof0 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=55531'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=55531'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-55531</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2000-03-23 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-09 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55531</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Occurs in fresh and brackish waters with salinity up to 13 ppt.  Maximum depth 50 m, with mature adults at 1-11 m.  Lives for about a year, spawns in May-August in northern Caspian, April-November in eastern Caspian.  Some individuals may spawn for the first time at 6-7 months.  Females probably lay eggs in 3 portions and die after spawning while males probably die 3-4 weeks alter (Ref. 59043).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27116&amp;speccode=55531'>McAllister, D.E. 1990 A working list of fishes of the world. Copies available from D.E. McAllister, Canadian Museum of Nature, P.O. Box 3443, Ottawa, Ontario K1P 6P4, Canada. 2661 p. plus 1270 p. Index. (Ref. 27116)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-55531</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2010-06-18 00:00:00</dcterms:created>
	<dcterms:modified>2010-06-18 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55531</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>This species is distinguished from its congeners by the following characters:  strongly depressed head; a tubercle between eyes; numerous, densely-set granules covering whole body except abdominal area; small tubercles especially on head; tubercles in dorsal row 26-29, ventral row 21-24, upper lateral row up to 20 (smaller on caudal peduncle), in lower one 10 and are tiny; chin barbel slightly compressed with thick base, longer than half eye diameter; D1 IV; origin of D2 behind anal origin; transverse rows of neuromasts on flank usually 20-22; a dark spot often in front of D2; sides with dark blotches and irregular dots (Ref. 59043).</dc:description>
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	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=55531'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-55531</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2000-03-23 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55531</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Eurasia:  Caspian eastern coast of Caspian Sea and Volga delta, up to Volgograd.</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27116&amp;speccode=55531'>McAllister, D.E. 1990 A working list of fishes of the world. Copies available from D.E. McAllister, Canadian Museum of Nature, P.O. Box 3443, Ottawa, Ontario K1P 6P4, Canada. 2661 p. plus 1270 p. Index. (Ref. 27116)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-55531</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>2000-03-23 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-09 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=55531&amp;StockCode=45939</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Females probably lay eggs in 2-3portions (Ref. 59043).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=55531'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-55531</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2000-03-23 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-09 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55531</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>6.6 cm TL (male/unsexed; (Ref. 50519)); max. reported age: 1 years (Ref. 59043)</dc:description>
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	<reference isbn='5-02-002872-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=50519&amp;speccode=55531'>Reshetnikov, Y.S. 2003 Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519)</reference>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=55531'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:rights></dc:rights>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-55531</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2000-03-23 00:00:00</dcterms:created>
	<dcterms:modified>2024-05-09 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=55531</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>demersal; freshwater; brackish; depth range 1 - 50 m (Ref. 59043)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='978-2-8399-0298-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=59043&amp;speccode=55531'>Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043)</reference></dataObject>
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