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	<dc:identifier>FB-5572</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Acanthuriformes</dwc:Order>
	<dwc:Family>Chaetodontidae</dwc:Family>
	<dwc:Genus>Chaetodon</dwc:Genus>
	<dwc:ScientificName>Chaetodon quadrimaculatus Gray, 1831</dwc:ScientificName>
	<commonName xml:lang='da'>Firplettet fanefisk</commonName>
	<commonName xml:lang='en'>Four-spot butterfly</commonName>
	<commonName xml:lang='en'>Fourspot butterflyfish</commonName>
	<commonName xml:lang='en'>Neonbanded butterflyfish</commonName>
	<commonName xml:lang='fj'>daliga ragutu</commonName>
	<commonName xml:lang=''>Lau-hau</commonName>
	<commonName xml:lang=''>Lau-lau</commonName>
	<commonName xml:lang='ja'>Shiten-chôchô-uo</commonName>
	<commonName xml:lang='sm'>Tifitifi-segasega</commonName>
	<commonName xml:lang='ty'>Paraharaha</commonName>
	<commonName xml:lang=''>Te ibaba</commonName>
	<commonName xml:lang=''>Te maninipapa</commonName>
	<commonName xml:lang=''>Tikorekore</commonName>
	<commonName xml:lang=''>四点蝴蝶鱼</commonName>
	<commonName xml:lang=''>四点蝶</commonName>
	<commonName xml:lang=''>四點蝴蝶魚</commonName>
	<synonym relationship='synonym'>Chaetodon quadromaculatus Gray, 1831</synonym>
	<synonym relationship='synonym'>Heterochaetodon lepidochaetodon quadrimaculatus (Gray, 1831)</synonym>
	<dcterms:created>1992-06-05</dcterms:created>
	<dcterms:modified>2014-10-13</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=5572'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=240&amp;speccode=5572'>Thresher, R.E.0 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=559&amp;speccode=5572'>Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino0 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=592&amp;speccode=5572'>Wass, R.C.0 An annotated checklist of the fishes of Samoa. Natl. Ocean. Atmos. Adminis. Tech. Rept., Natl. Mar. Fish. Serv., Spec. Sci. Rept. Fish. (781). (Ref. 592)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1602&amp;speccode=5572'>Myers, R.F.0 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3804&amp;speccode=5572'>Dalzell, P., S.R. Lindsay and H. Patiale0 Fisheries resources survey of the Island of Niue. Tech. Doc. Inshore Fish. Res. Proj. S. Pac. Comm 3. A report prepared in conjunction with the South Pacific Commission Inshore Fisheries Research Project, and the FAO South Pacific Aquaculture Development Project for the Government of Niue, July 1990. (Ref. 3804)</reference>
	<reference isbn='0-915180-07-3 or 0-915180-29-4?' url='http://www.fishbase.org/references/FBRefSummary.php?id=3921&amp;speccode=5572'>Randall, J.E.0 Guide to Hawaiian reef fishes. Harrowood Books, Newtown Square, PA 19073, USA. 74 p. (Ref. 3921)</reference>
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	<reference isbn='3-89356-170-6.' url='http://www.fishbase.org/references/FBRefSummary.php?id=9137&amp;speccode=5572'>Masuda, H. and G.R. Allen0 Meeresfische der Welt - Groß-Indopazifische Region. Tetra Verlag, Herrenteich, Melle. 528 p. (Ref. 9137)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=5572'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<dc:description>Adults are found almost exclusively on exposed seaward reefs (Ref. 205).  Benthopelagic (Ref. 58302).  They feed mainly on polyps of Pocillopora.  Solitary or in pairs.  Oviparous (Ref. 205), monogamous (Ref. 52884).  Form pairs during breeding (Ref. 205).</dc:description>
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	<dc:title>Importance</dc:title>
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	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5572</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description>fisheries: commercial; aquarium: commercial</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5358&amp;speccode=5572'>Miyasaka, A. 1993 A database on scientific and common names of fishes exported from Hawaii. The information was derived from the above mentioned database. A printout of the names is also available from the State of Hawaii, Department of Land and Natural Resources, 1151 Punchbowl Street, Honolulu, Hawaii. (Ref. 5358)</reference></dataObject><dataObject>
	<dc:identifier>FB-LifeCycle-5572</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1992-06-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-20 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=5572&amp;StockCode=5862</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Distinct pairing (Ref. 205). Monogamous mating is observed as obligate, genetic and social (Ref. 52884).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=240&amp;speccode=5572'>Thresher, R.E. 1984 Reproduction in reef fishes. T.F.H. Publications, Inc. Ltd., Neptune City, New Jersey. 399 p. (Ref. 240)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-5572</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1997-03-25 00:00:00</dcterms:created>
	<dcterms:modified>2001-11-20 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>quadrimaculatus</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Found almost exclusively on exposed seaward reefs. Feeds mainly on polyps of Pocillopora (Ref. 1602).</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='0-915180-07-3 or 0-915180-29-4?' url='http://www.fishbase.org/references/FBRefSummary.php?id=3921&amp;speccode=5572'>Randall, J.E. 1985 Guide to Hawaiian reef fishes. Harrowood Books, Newtown Square, PA 19073, USA. 74 p. (Ref. 3921)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-5572</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1992-06-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-20 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5572</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>16.0 cm TL (male/unsexed; (Ref. 9710))</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='0-00-219974-2 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=9710&amp;speccode=5572'>Lieske, E. and R. Myers 1994 Collins Pocket Guide. Coral reef fishes. Indo-Pacific &amp; Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-5572</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
	<mimeType>text/html</mimeType>
	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1992-06-05 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-20 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=5572</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses</subject>
	<dc:description>reef-associated; non-migratory (Ref. 52884); marine; depth range 0 - 43 m (Ref. 89972), usually 2 - 15 m (Ref. 1602)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52884&amp;speccode=5572'>Whiteman, E.A. and I.M. Côté 2004 Monogamy in marine fishes. Biol. Rev. 79:351-375. (Ref. 52884)</reference>
	<reference isbn='2-915654-09-3' url='http://www.fishbase.org/references/FBRefSummary.php?id=89972&amp;speccode=5572'>Bacchet, P., T. Zysman and Y. Lefèvre 2006 Guide des poissons de Tahiti et ses îles. Tahiti (Polynésie Francaise): Éditions Au Vent des Îles. 608 p. (Ref. 89972)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1602&amp;speccode=5572'>Myers, R.F. 1991 Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602)</reference></dataObject>
</taxon>
</response>