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<taxon>
	<dc:identifier>FB-62419</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62419</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Nannacara</dwc:Genus>
	<dwc:ScientificName>Nannacara quadrispinae Staeck &amp; Schindler, 2004</dwc:ScientificName>
	<commonName xml:lang=''>Kolonto</commonName>
	<commonName xml:lang='my'>Hnit Yaung Phyar Kyo Warr</commonName>
	<commonName xml:lang='en'>Spotted goatfish</commonName>
	<commonName xml:lang='en'>Twotone goatfish</commonName>
	<commonName xml:lang='en'>Two-tone goatfish</commonName>
	<commonName xml:lang='sd'>Adrakh</commonName>
	<commonName xml:lang='sd'>Chakori</commonName>
	<synonym relationship='synonym'>Upeneoides guttatus Day, 1868</synonym>
	<synonym relationship='synonym'>Upeneus crosnieri Fourmanoir &amp; Guézé, 1967</synonym>
	<synonym relationship='misapplied name'>Upeneus pori (non Ben-Tuvia &amp; Golani, 1989)</synonym>
	<synonym relationship='misapplied name'>Upeneus taeniopterus (non Cuvier, 1829)</synonym>
	<dcterms:created>2005-12-01</dcterms:created>
	<dcterms:modified>2022-01-31</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36648&amp;speccode=62425'>Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43139&amp;speccode=62425'>Kim, B.-J. and K. Nakaya0 &lt;i&gt;Upeneus australiae&lt;/i&gt;, a new goatfish (Mullidae:Perciformes) from Australia. Ichthyol. Res. 49(2):128-132. (Ref. 43139)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48643&amp;speccode=62425'>Khalaf, M.A. and A.M. Disi0 Fishes of the Gulf of Aqaba. Marine Science Station, Aqaba, Jordan. 252 p. (Ref. 48643)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=54621&amp;speccode=62425'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of May 2005. Database dump. Catalog databases as made available to FishBase in May 2005. (Ref. 54621)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55787&amp;speccode=62425'>Kulbicki, M., N. Guillemot and M. Amand0 A general approach to length-weight relationships for New Caledonian lagoon fishes. Cybium 29(3):235-252. (Ref. 55787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55797&amp;speccode=62425'>Kulbicki, M., Y.-M. Bozec, P. Labrosse, Y. Letourneur, G. Mou-Tham and L. Wantiez0 Diet composition of carnivorous fishes from coral reef lagoons of New Caledonia. Aquat. Living Resour. 18:231-250. (Ref. 55797)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58005&amp;speccode=62425'>Randall, J.E. and M. Kulbicki0 A review of the goatfishes of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Perciformes: Mullidae) from New Caledonia and the Chesterfield Bank, with a new species and four new records. Zool. Studies 45(3):298-307. (Ref. 58005)</reference>
	<reference isbn='978-2-7592-0045-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=81519&amp;speccode=62425'>Taquet, M. and A. Diringer0 Poissons de l'Océan Indien et de la mer Rouge. Versailles (France): Éditions Quæ. 527 p. (Ref. 81519)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83903&amp;speccode=62425'>Uiblein, F. and P.C. Heemstra0 A taxonomic review of the Western Indian Ocean goatfishes of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Family Mullidae), with descriptions of four new species. Smithiana 11:35-71. (Ref. 83903)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=84159&amp;speccode=62425'>Golani, D. and S. V. Bogorodsky0 The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa 2463:1-135. (Ref. 84159)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=86942&amp;speccode=62425'>Fricke, R., M. Kulbicki and L. Wantiez0 Checklist of the fishes of New Caledonia, and their distribution in the Southwest Pacific Ocean (Pisces). Stuttgarter Beiträge zur Naturkunde A, Neue Serie 4:341-463. (Ref. 86942)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87596&amp;speccode=62425'>Rajan, P.T., C.R. Sreeraj and T. Immanuel0 Fish fauna of coral reef, mangrove, freshwater, offshore and seagrass beds of Andaman and Nicobar Islands. Zoological Survey of India, Andaman and Nicobar Regional Centre, Haddo, Port Blair. (Ref. 87596)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=94148&amp;speccode=62425'>Uiblein, F. and M. Lisher0 A new goatfish of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Mullidae) from Angoche, northern Mozambique. Zootaxa 3717(1):85-95. (Ref. 94148)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=98481&amp;speccode=62425'>Uiblein, F. and D.C. Gledhill0 A new goatfish of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Mullidae) from Australia and Vanuatu, with inter- and intraspecific comparisons. Mar. Biol. Res. 25(11):475-491. (Ref. 98481)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=99978&amp;speccode=62425'>Bogorodsky, S.V., T.J. Alpermann, A.O. Mal and M.H. Gabr0 Survey of the demersal fishes from southern Saudi Arabia, with five new records for the Red Sea. Zootaxa 3852(4):401-437. (Ref. 99978)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101194&amp;speccode=62425'>Patzner, R.A.0 Reproductive strategies of fish. pp. 311-350.  In  Rocha, M.J., A. Arukwe and B.G. Kapoor (eds). Fish reproduction: cytology, biology and ecology. Science Publisher, Inc. Oxford. 631 p. (Ref. 101194)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=102936&amp;speccode=62425'>Uiblein, F. and W.T. White0 A new goatfish of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Mullidae) from Lombok, Indonesia and first verified record of &lt;i&gt;U. asymmetricus&lt;/i&gt; for the Indian Ocean. Zootaxa 3980(1):51-66. (Ref. 102936)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=110201&amp;speccode=62425'>Uiblein, F., G. Gouws, D.C. Gledhill and K. Stone0 Just off the beach: intrageneric distinctiveness of the bandtail goatfish &lt;i&gt;Upeneus taeniopterus&lt;/i&gt; (Mullidae) based on a comprehensive alpha-taxonomy and barcoding approach. Mar. Biol. Res. 12(7):675-694. (Ref. 110201)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114767&amp;speccode=62425'>Psomadakis, P.N., H.B. Osmany and M. Moazzam0 Field identification guide to the living marine resources of Pakistan. FAO Species Identification Guide for Fishery Purposes, 386p. (Ref. 114767)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=116177&amp;speccode=62425'>Uiblein, F., D.C. Gledhill and T. Peristiwady0 Two new goatfishes of the genus &lt;i&gt;Upenus&lt;/i&gt; (Mullidae) from Australia and Indonesia. Zootaxa 4318(1):295-311. (Ref. 116177)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122869&amp;speccode=62425'>Psomadakis, P., H. Thein, B.C. Russell and M.T. Tun0 Field identification guide to the living marine resources of Myanmar. FAO species identification guide for fishery purposes. Food and Agriculture Organization of the United Nations, and Department of Fisheries, Ministry of Agriculture, Livestock and Irrigation, Republic of the Union of Myanmar, Rome: i-xvii + 1-694, pls. 1-58. (Ref. 122869)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123119&amp;speccode=62425'>Uiblein, F., G. Gouws, M. Lisher and B.S. Malauene0 &lt;i&gt;Upeneus floros&lt;/i&gt;, a new goatfish from South Africa and Mozambique, with updated taxonomic accounts for &lt;i&gt;U. guttatus&lt;/i&gt; and &lt;i&gt;U. pori&lt;/i&gt; and a key to Western Indian Ocean Upeneus species (Mullidae). Zootaxa 4834(4):523-555. (Ref. 123119)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123517&amp;speccode=62425'>Motomura, H., M. Yamashita, M. Itou, Y. Yaraguchi and Y. Iwatsuki0 First records of the Two-tone goatsh,&lt;i&gt;Upeneus guttatus&lt;/i&gt; from Japan, and comparisons with &lt;i&gt;U. japonicus&lt;/i&gt; (Perciformes: Mullidae). Species Diversity 17:7-14. (Ref. 123517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=126597&amp;speccode=62425'>Sabrah, M.M., R.A. Heneish, M.E. Alwany and Md.I. Ahmad0 Sexual maturity, spawning activity, sex ratio and fecundity of two Mullidae species dwelling the Gulf of Suez, Red Sea. The Egyptian Journal of Aquatic Research 43(1):83-91. (Ref. 126597)</reference><dataObject>
	<dc:identifier>FB-DiagnosticDescr-62425</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2010-04-12 00:00:00</dcterms:created>
	<dcterms:modified>2021-01-28 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62425</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>This species is distinguished by the following characters:  D VII + 9; pectoral fins 12-15; gill rakers 5-8 + 16-19 = 23-26.  Measurements as %SL for adults: body depth at first dorsal fin origin 21-26 and at anus 18-22; caudal-peduncle depth 9.3-11; maximum head depth 18-22 and through eye 15-18; head length 26-29; snout length 9.4-12; orbit length 5.9-8.5; upper jaw length 9.5-12;  barbel length 16-20; caudal-fin length 27-31; anal-fin height 15-19; pelvic-fin length 19-22; pectoral-fin length 19-22; first dorsal-fin height 20-25, at least one of the first 3 spines disproportionally higher; second dorsal-fin height 14-18; total oblique bars on caudal fin 7-16; upper caudal-fin lobe with 5-6 (rarely 4) reddish bars with 4-5 (rarely 3) bars distally from fork and one bar close to fin lobe base, of similar width or narrower than the pale interspaces between bars; there are 2-10 often faint, rather short and/or irregularly arranged red bars on ventral margin of lower caudal-fin lobe, sometimes extending to dorsal half of lobe or connecting to a red stripe which covers the lobe dorsally to two-thirds of its width at maximum; the caudal-fin lobe bars and stripe fade away after death and only traces retained in preserved fish; when fresh, barbels yellow rarely white; body colour may be variable, often white or rose below lateral line, covered by red pigmentation above lateral line which may also reach down ventrally and to head, sometimes forming red patches or blotches; body raely with a faint red or red-orange mid-lateral stripe from behind eye to caudal-fin base; when preserved, the body is pale brown and not dorsally darkened (Ref. 83903).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123119&amp;speccode=62425'>Uiblein, F., G. Gouws, M. Lisher and B.S. Malauene 2020 &lt;i&gt;Upeneus floros&lt;/i&gt;, a new goatfish from South Africa and Mozambique, with updated taxonomic accounts for &lt;i&gt;U. guttatus&lt;/i&gt; and &lt;i&gt;U. pori&lt;/i&gt; and a key to Western Indian Ocean Upeneus species (Mullidae). Zootaxa 4834(4):523-555. (Ref. 123119)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-62425</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2010-04-12 00:00:00</dcterms:created>
	<dcterms:modified>2021-01-28 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62425</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Dorsal spines (total): 7; Dorsal soft rays (total): 9</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123119&amp;speccode=62425'>Uiblein, F., G. Gouws, M. Lisher and B.S. Malauene 2020 &lt;i&gt;Upeneus floros&lt;/i&gt;, a new goatfish from South Africa and Mozambique, with updated taxonomic accounts for &lt;i&gt;U. guttatus&lt;/i&gt; and &lt;i&gt;U. pori&lt;/i&gt; and a key to Western Indian Ocean Upeneus species (Mullidae). Zootaxa 4834(4):523-555. (Ref. 123119)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-62425</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2005-12-06 00:00:00</dcterms:created>
	<dcterms:modified>2025-03-05 00:00:00</dcterms:modified>
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	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62425</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Indo-West Pacific: from S Mozambique and the Red Sea to Japan, N Australia and New Caledonia.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123119&amp;speccode=62425'>Uiblein, F., G. Gouws, M. Lisher and B.S. Malauene 2020 &lt;i&gt;Upeneus floros&lt;/i&gt;, a new goatfish from South Africa and Mozambique, with updated taxonomic accounts for &lt;i&gt;U. guttatus&lt;/i&gt; and &lt;i&gt;U. pori&lt;/i&gt; and a key to Western Indian Ocean Upeneus species (Mullidae). Zootaxa 4834(4):523-555. (Ref. 123119)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-62425</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2005-12-06 00:00:00</dcterms:created>
	<dcterms:modified>2021-01-28 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62425</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>16.0 cm SL (male/unsexed; (Ref. 83903))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83903&amp;speccode=62425'>Uiblein, F. and P.C. Heemstra 2010 A taxonomic review of the Western Indian Ocean goatfishes of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Family Mullidae), with descriptions of four new species. Smithiana 11:35-71. (Ref. 83903)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-62425</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2005-12-06 00:00:00</dcterms:created>
	<dcterms:modified>2021-01-28 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=62425</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>demersal; marine; depth range 8 - 165 m (Ref. 98481)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=98481&amp;speccode=62425'>Uiblein, F. and D.C. Gledhill 2015 A new goatfish of the genus &lt;i&gt;Upeneus&lt;/i&gt; (Mullidae) from Australia and Vanuatu, with inter- and intraspecific comparisons. Mar. Biol. Res. 25(11):475-491. (Ref. 98481)</reference></dataObject>
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