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	<dc:identifier>FB-6371</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Serranoidei</dwc:Order>
	<dwc:Family>Epinephelidae</dwc:Family>
	<dwc:Genus>Hyporthodus</dwc:Genus>
	<dwc:ScientificName>Hyporthodus quernus (Seale, 1901)</dwc:ScientificName>
	<commonName xml:lang='ar'>لُؤز هاوايي</commonName>
	<commonName xml:lang='en'>Grouper</commonName>
	<commonName xml:lang='en'>Hawaiian black grouper</commonName>
	<commonName xml:lang='en'>Hawaiian grouper</commonName>
	<commonName xml:lang='en'>Hawaiian sea bass</commonName>
	<commonName xml:lang='en'>Sea bass</commonName>
	<commonName xml:lang='en'>Seale's grouper</commonName>
	<commonName xml:lang='fr'>Mérou hawaiien</commonName>
	<commonName xml:lang=''>Hapu'u</commonName>
	<commonName xml:lang=''>Hapuu puu</commonName>
	<commonName xml:lang=''>Hapu'upu'u</commonName>
	<commonName xml:lang='he'>Lokos</commonName>
	<commonName xml:lang='es'>Mero hawaiano</commonName>
	<commonName xml:lang=''>夏威夷石斑魚</commonName>
	<commonName xml:lang=''>夏威夷石斑鱼</commonName>
	<synonym relationship='synonym'>Epinephelus quernus Seale, 1901</synonym>
	<dcterms:created>1992-08-20</dcterms:created>
	<dcterms:modified>2020-11-06</dcterms:modified>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=6371'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=583&amp;speccode=6371'>Tinker, S.W.0 Fishes of Hawaii, a handbook of the marine fishes of Hawaii and the Central Pacific Ocean. Hawaiian Service Inc., Honolulu. 568 p. (Ref. 583)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1830&amp;speccode=6371'>Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4787&amp;speccode=6371'>Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4887&amp;speccode=6371'>Honebrink, R.0 Fishing in Hawaii: a student manual. Education Program, Division of Aquatic Resources, Honolulu, Hawaii. 79 p. (Ref. 4887)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6371'>Heemstra, P.C. and J.E. Randall0 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5358&amp;speccode=6371'>Miyasaka, A.0 A database on scientific and common names of fishes exported from Hawaii. The information was derived from the above mentioned database. A printout of the names is also available from the State of Hawaii, Department of Land and Natural Resources, 1151 Punchbowl Street, Honolulu, Hawaii. (Ref. 5358)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7277&amp;speccode=6371'>Seki, M.P.0 Serranidae. p. 82-83. In R.N. Uchida and J.H. Uchiyama (eds.) Fishery Atlas of the Northwestern Hawaiian Islands. NOAA Tech. Rep. NMFS 38. (Ref. 7277)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28620&amp;speccode=6371'>Randall, J.E.0 Randall's underwater photos. Collection of almost 2,000 underwater photos (slides). Unpublished. (Ref. 28620)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=6371'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31667&amp;speccode=6371'>Anonymous0 Fish collection database of the Bernice P. Bishop Museum (BPBM). Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35722&amp;speccode=6371'>Uchida, R.N. and J.H. Uchiyama0 Fishery Atlas of the Northwestern Hawaiian Islands. NOAA Tech. Rep. NMFS 38,142 p. (Ref. 35722)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=6371'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40637&amp;speccode=6371'>IGFA0 Database of IGFA angling records until 2001. IGFA, Fort Lauderdale, USA. (Ref. 40637)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=6371'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50505&amp;speccode=6371'>Humphreys, R.L. Jr., D.T. Tagami and M.P. Seki0 Seamount fishery resources within the southern Emperor-Northern Hawaiian Ridge. Second symposium on resources investigations in the Northwestern Hawaiian Islands (Honolulu, Hawaii, 25-27 May 1983), R.W. Grigg and K.Y. Tanoue, vol 1, pp. 283-327. (Ref. 50505)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50612&amp;speccode=6371'>Uchida, R.N. and D.T. Tagami0 Groundfish fisheries and research in the vicinity of seamounts in the north Pacific ocean. Marine Fisheries Review 46(2):1-17. (Ref. 50612)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51159&amp;speccode=6371'>DeMartini, E.E. and A.M. Friedlander0 Spatial patterns of endemism in shallow-water reef fish populations of the northwestern Hawaiian islands. Mar. Ecol. Prog. Ser. 271:281-296. (Ref. 51159)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=6371'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58302&amp;speccode=6371'>Mundy, B.C.0 Checklist of the fishes of the Hawaiian Archipelago. Bishop Mus. Bull. Zool. (6):1-704. (Ref. 58302)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83414&amp;speccode=6371'>Craig, M.T. and P.A. Hastings0 A molecular phylogeny of the groupers of the subfamily Epinephelinae (Serranidae) with revised classification of the epinephelini. Ichthyol. Res. 54:1-17. (Ref. 83414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='9781466506022' url='http://www.fishbase.org/references/FBRefSummary.php?id=89707&amp;speccode=6371'>Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=111088&amp;speccode=6371'>Nichols, R.S. and E.E. DeMartini0 Preliminary estimates of age and growth for the endemic Hawaiian grouper Hapu’upu’u, &lt;i&gt;Epinephelus quernus&lt;/i&gt;, F. Serranidae). Pacific Islands Fish. Sci. Cent., Natl. Mar. Fish. Serv., NOAA, Honolulu, HI 96822-2396. Pacific Islands Fish. Sci. Cent. Admin. Rep. H-08-06, 19 p. (Ref. 111088)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=6371'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=122680&amp;speccode=6371'>State of hawaii0 SWAP species fact sheets. https://dlnr.hawaii.gov/wildlife/hswap/fact-sheets/ [Accessed 13/08/2020]. (Ref. 122680)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=6371'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='http://www.fishbase.us/collaborators/CollaboratorSummary.php?id=823' logoURL='' role='photographer'> FAO</agent>
	<dcterms:created>1996-05-26</dcterms:created>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1992-08-20 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Occurs in deep waters (Ref. 7364).  Benthic and benthopelagic (Ref. 58302).  Feeds mainly on fish and crustaceans (Ref. 89707). A good fish to eat, fleshy, eaten any way except raw (Ref. 7364).  Minimum depth from Ref. 122680.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83414&amp;speccode=6371'>Craig, M.T. and P.A. Hastings 2007 A molecular phylogeny of the groupers of the subfamily Epinephelinae (Serranidae) with revised classification of the epinephelini. Ichthyol. Res. 54:1-17. (Ref. 83414)</reference></dataObject><dataObject>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Distinguished by the following characteristics:  juveniles greyish brown to dark brown body color with 8 vertical series of white spots on body; adults dark brown with less distinct vertical series of white spots and obscured by numerous additional pale spots and blotches of variable size; fins mostly unspotted; depth of body contained 2.3-2.7 times in SL, width 2.0-2.5 times in depth; head length 2.3-2.5 times in SL; convex interorbital area, width subequal to eye diameter for fish 10-20cm SL; subangular preopercle, 3-5 enlarged serrae at angle; fleshy ventral edge of preopercle, without serrae; smooth edge of interopercle and subopercle or with a few serrae; convex upper edge of operculum; ovate and enlarged posterior nostrils of adults, larger than anterior nostrils (Ref. 89707).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6371'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
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	<dc:description>Dorsal spines (total): 11; Dorsal soft rays (total): 14 - 15;  Analspines: 3; Analsoft rays: 9</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6371'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<dc:description>Eastern Central Pacific:  known only from the Hawaiian Islands and Johnston Island.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6371'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<dc:description>fisheries: commercial; gamefish: yes; aquarium: commercial</dc:description>
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	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=6371'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4887&amp;speccode=6371'>Honebrink, R. 1990 Fishing in Hawaii: a student manual. Education Program, Division of Aquatic Resources, Honolulu, Hawaii. 79 p. (Ref. 4887)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5358&amp;speccode=6371'>Miyasaka, A. 1993 A database on scientific and common names of fishes exported from Hawaii. The information was derived from the above mentioned database. A printout of the names is also available from the State of Hawaii, Department of Land and Natural Resources, 1151 Punchbowl Street, Honolulu, Hawaii. (Ref. 5358)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4887&amp;speccode=6371'>Honebrink, R. 1990 Fishing in Hawaii: a student manual. Education Program, Division of Aquatic Resources, Honolulu, Hawaii. 79 p. (Ref. 4887)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4887&amp;speccode=6371'>Honebrink, R. 1990 Fishing in Hawaii: a student manual. Education Program, Division of Aquatic Resources, Honolulu, Hawaii. 79 p. (Ref. 4887)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4887&amp;speccode=6371'>Honebrink, R. 1990 Fishing in Hawaii: a student manual. Education Program, Division of Aquatic Resources, Honolulu, Hawaii. 79 p. (Ref. 4887)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:description>benthopelagic; marine; depth range 5 - 380 m (Ref. 5222)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6371'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject>
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