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	<dc:identifier>FB-6460</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Perciformes/Serranoidei</dwc:Order>
	<dwc:Family>Epinephelidae</dwc:Family>
	<dwc:Genus>Epinephelus</dwc:Genus>
	<dwc:ScientificName>Epinephelus bontoides (Bleeker, 1855)</dwc:ScientificName>
	<commonName xml:lang='en'>Dusky rock cod</commonName>
	<commonName xml:lang='en'>Palemargin grouper</commonName>
	<commonName xml:lang='fr'>Mérou bord clair</commonName>
	<commonName xml:lang='he'>Lokos</commonName>
	<commonName xml:lang='tl'>Lapu-lapu</commonName>
	<commonName xml:lang='es'>Mero de márgenes pálidos</commonName>
	<commonName xml:lang=''>Lapu-lapu</commonName>
	<commonName xml:lang=''>点列石斑鱼</commonName>
	<commonName xml:lang=''>石斑</commonName>
	<commonName xml:lang=''>过鱼</commonName>
	<commonName xml:lang=''>鲙</commonName>
	<commonName xml:lang=''>點列石斑魚</commonName>
	<synonym relationship='synonym'>Serranus bontoides Bleeker, 1855</synonym>
	<dcterms:created>1992-10-13</dcterms:created>
	<dcterms:modified>2013-09-02</dcterms:modified>
	<reference isbn='92-5-103623-3.' url='http://www.fishbase.org/references/FBRefSummary.php?id=171&amp;speccode=6460'>Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=280&amp;speccode=6460'>Herre, A.W.C.T.0 Check list of Philippine fishes. Res. Rep. U.S. Fish Wild. Serv. (20):1-977. (Ref. 280)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=395&amp;speccode=6460'>Bleeker, P.0 Zesde bijdrage tot de kennis der ichthyologische fauna van Amboina. Natuurkundig Tijdschrift voor Nederlandsch Indië 8(3):391-434. (Ref. 395)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1830&amp;speccode=6460'>Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2857&amp;speccode=6460'>Herre, A.W.C.T. and A.F. Umali0 English and local common names of Philippine fishes. U. S. Dept. of Interior and Fish and Wildl. Serv. Circular No. 14, U. S. Gov't Printing Office, Washington. 128 p. (Ref. 2857)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4787&amp;speccode=6460'>Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall0 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6192&amp;speccode=6460'>Kailola, P.J.0 The fishes of Papua New Guinea: a revised and annotated checklist. Vol. II Scorpaenidae to Callionymidae. Research Bulletin No. 41, Research Section, Dept. of Fisheries and Marine Resources, Papua New Guinea. (Ref. 6192)</reference>
	<reference isbn='979-511-058-6.' url='http://www.fishbase.org/references/FBRefSummary.php?id=8631&amp;speccode=6460'>Kuiter, R.H.0 Tropical reef-fishes of the western Pacific Indonesia and adjacent waters. Gramedia Pustaka Utama, Jakarta. 314 p. (Ref. 8631)</reference>
	<reference isbn='0-00-219974-2 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=9710&amp;speccode=6460'>Lieske, E. and R. Myers0 Collins Pocket Guide. Coral reef fishes. Indo-Pacific &amp; Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12846&amp;speccode=6460'>Nguyen, H.P., P. Le Trong, N.T. Nguyen, P.D. Nguyen, N. Do Thi Nhu and V.L. Nguyen0 Checklist of marine fishes in Vietnam. Vol. 3. Order Perciformes, Suborder Percoidei, and Suborder Echeneoidei. Science and Technics Publishing House, Vietnam. (Ref. 12846)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28618&amp;speccode=6460'>Randall, J.E.0 Randall's tank photos. Collection of 10,000 large-format photos (slides) of dead fishes. Unpublished. (Ref. 28618)</reference>
	<reference isbn='971-8722-02-5.' url='http://www.fishbase.org/references/FBRefSummary.php?id=31411&amp;speccode=6460'>Ganaden, S.R. and F. Lavapie-Gonzales0 Common and local names of marine fishes of the Philippines. Bureau of Fisheries and Aquatic Resources, Philippines. 385 p. (Ref. 31411)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=6460'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31667&amp;speccode=6460'>Anonymous0 Fish collection database of the Bernice P. Bishop Museum (BPBM). Bishop Museum, 1525 Bernice Street, Honolulu, Hawai`I, 96817-0916 USA. (Ref. 31667)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=6460'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36648&amp;speccode=6460'>Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=6460'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=39231&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall0 Serranidae. p. 2442-2547. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 4. Bony fishes part 2 (Mugilidae to Carangidae). Rome, FAO. 2069-2790 p. (Ref. 39231)</reference>
	<reference isbn='1-881173-54-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=41464&amp;speccode=6460'>Allen, G.R.0 Reef fishes of Milne Bay Province, Papua New Guinea. In T. Werner and G. Allen (eds). A rapid biodiversity assessment of the coral reefs of Milne Bay Province, Papua New Guinea. RAP Working Papers 11, Conservation International, Washington, D.C. (Ref. 41464)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47567&amp;speccode=6460'>Allen, G.R. and M. Adrim0 Coral reef fishes of Indonesia. Zool. Stud. 42(1):1-72. (Ref. 47567)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=47843&amp;speccode=6460'>Huang, Z.0 Marine species and their distribution in China's seas. p. 404-463. Vertebrata. Smithsonian Institution, Florida, USA. 598 p. (Ref. 47843)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48497&amp;speccode=6460'>Hardy, J.D. Jr. (comp.)0 Coral reef fish species. NOAANational Oceanographic Data Center. NODC Coral Reef Data and Information Management System. USA. 537 p. (Ref. 48497)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=48635&amp;speccode=6460'>Kuiter, R.H. and T. Tonozuka0 Pictorial guide to Indonesian reef fishes. Part 1. Eels- Snappers, Muraenidae - Lutjanidae. Zoonetics, Australia. 1-302. (Ref. 48635)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=6460'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='9781466506022' url='http://www.fishbase.org/references/FBRefSummary.php?id=89707&amp;speccode=6460'>Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707)</reference>
	<reference isbn='9780987260000' url='http://www.fishbase.org/references/FBRefSummary.php?id=90102&amp;speccode=6460'>Allen, G.R. and M.V. Erdmann0 Reef fishes of the East Indies. Perth, Australia: Universitiy of Hawai'i Press, Volumes I-III. Tropical Reef Research. (Ref. 90102)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=121724&amp;speccode=6460'>Songco, A.0 Personal communication. Tubbataha species list. Excel file "MyReport_Species_21May2019". (Ref. 121724)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130287&amp;speccode=6460'>Balisco, R.A.T., V.S. Ticzon, B.R. Samaniego, W.-C. Huang, B.J. Gonzales and T.-Y. Liao0 Marine fishes of Palawan, Philippines: species diversity, new records, and conservation status. Regional Studies in Marine Science 60(2023) 102825:1-10. (Ref. 130287)</reference><dataObject>
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	<dcterms:created>1996-03-01</dcterms:created>
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	<dc:description>Sao Wisata Resort Locality: Maumere, Flores, Indonesia.</dc:description>
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	<location>Locality: Maumere, Flores, Indonesia.</location></dataObject><dataObject>
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	<dcterms:created>1996-05-25</dcterms:created>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>1992-10-13 00:00:00</dcterms:created>
	<dcterms:modified>2013-09-02 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>Found on mud or rocky or cobble bottoms (Ref. 089707).  The species appears to be one of the rarest groupers in the Indo-Pacific region, nothing has been published on its biology. Solitary (Ref 90102).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
	<dcterms:created>1994-06-10 00:00:00</dcterms:created>
	<dcterms:modified>2013-09-02 00:00:00</dcterms:modified>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Characterized by grey brown color with scattered reddish brown to black spots; narrow pale margins on dark grey fins; body scales ctenoid, except cycloid scales dorsally and ventrally; body with auxiliary scales; greatest depth of body 2.8-3.2 in SL; rounded caudal fin; pelvic fins, 1.9-2.3 in head length (Ref. 90102); head length 2.3-2.6 times in SL; interorbital area flat to slightly convex, almost straight dorsal head profile; rounded preopercle, finely serrate, ventral serrae slightly enlarged;  straight upper edge of operculum; maxilla reaches past rear edge of eye; 2 or 3 rows of teeth on midlateral part of lower jaw (Ref. 089707).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<agent homepage='https://sites.google.com/site/itsaboutfish/home#' logoURL='' role='recorder'>Rodolfo B. Reyes</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Dorsal spines (total): 11; Dorsal soft rays (total): 16 - 17;  Analspines: 3; Analsoft rays: 8</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<dc:description>Western Pacific:  Vietnam, Indonesia, Philippines, Taiwan, Solomon Islands (Ref. 4787) and New Britain.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5222&amp;speccode=6460'>Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)</reference></dataObject><dataObject>
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	<reference isbn='0-00-219974-2 (pbk.).' url='http://www.fishbase.org/references/FBRefSummary.php?id=9710&amp;speccode=6460'>Lieske, E. and R. Myers 1994 Collins Pocket Guide. Coral reef fishes. Indo-Pacific &amp; Caribbean including the Red Sea. Haper Collins Publishers, 400 p. (Ref. 9710)</reference></dataObject>
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