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	<dc:identifier>FB-66587</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66587</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Siluriformes</dwc:Order>
	<dwc:Family>Loricariidae</dwc:Family>
	<dwc:Genus>Harttiella</dwc:Genus>
	<dwc:ScientificName>Harttiella intermedia Covain &amp; Fisch-Muller, 2012</dwc:ScientificName>
	<synonym relationship='synonym'>Panaqolus koko Fisch-Muller &amp; Covain, 2012</synonym>
	<dcterms:created>2012-06-08</dcterms:created>
	<dcterms:modified>2012-06-08</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain0 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=116654&amp;speccode=66596'>Saks, L.0 Estonian Common Names, Rank 2. Personal communication (Ref. 116654)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=117193&amp;speccode=66596'>Lujan, N.K., C.A. Cramer, R. Covain, S. Fisch-Muller and H. López-Fernández0 Multilocus molecular phylogeny of the ornamental wood-eating catfishes (Siluriformes, Loricariidae, &lt;i&gt;Panaqolus&lt;/i&gt; and &lt;i&gt;Panaque&lt;/i&gt; reveals underscribed diversity and parapatric clades. Mol. Phy. Evol. 109: 321-336. (Ref. 117193)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2018-02-26 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>Found in main river channel on a stony substrate at two meters depth (Ref. 90183).  Collected together with the hypostomins Hemiancistrus medians, Peckoltia otali and Pseudancistrus barbatus, and with the loricariins Harttia guianensis, Loricaria cataphracta and Rineloricaria stewarti (Ref. 90183).  The gut contents of one specimen consisted mostly of intact sponge fragments and spicules (Ref. 117193).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain 2012 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2012-12-27 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>This species is distinguished by its large and almost spoon-shaped teeth characteristic of Panaqolus but bifid instead of most generally unicuspid in congeners, and characterized by its specific barcode sequence (JF747003).  Differs also from all other congeners except Panaqolus dentex and P. nocturnus by having a uniformly blackish-brown coloration (vs. banded pattern of coloration in P. purusiensis, P. gnomus, P. maccus, and P. changae; or spotted pattern of coloration in P. albomaculatus).  It can be separated from Panaqolus dentex by having dark pigment on membrane and branched rays of all fins, as well as a smaller interorbital width (29.4-33.2% HL, mean 31.6, vs. 38.7), a shorter pectoral spine (29.8-31.3% HL, mean 30.3, vs. 34.8) and a greater caudal peduncle depth (12.0-13.4% SL, mean 12.9, vs. 10.8).  It is readily distinguished from Panaqolus nocturnus by having larger eye (orbit length 18.8-20.8% HL vs. 13.7-15.9) (Ref. 90183).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain 2012 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2012-12-27 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>Dorsal spines (total): 2; Dorsal soft rays (total): 7;  Analspines: 1; Analsoft rays: 5</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain 2012 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>South America:  upper Maroni River basin.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain 2012 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2018-02-26 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>9.0 cm SL (male/unsexed; (Ref. 90183))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90183&amp;speccode=66596'>Fisch-Muller, S., J.I. Montoya-Burgos, P.-Y. Le Bail and R. Covain 2012 Diversity of the Ancistrini (Siluriformes: Locariidae) from the Guianas: the &lt;i&gt;Panaque&lt;/i&gt; group, a molecular appraisal with description of new species. Cybium 36(1):163-193. (Ref. 90183)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-66596</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Armi G. Torres</agent>
	<dcterms:created>2012-06-13 00:00:00</dcterms:created>
	<dcterms:modified>2018-02-26 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=66596</dc:source>
	<dc:description>demersal; freshwater</dc:description>
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