FB-7360 http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 Animalia Chordata Teleostei Perciformes/Serranoidei Epinephelidae Epinephelus Epinephelus radiatus (Day, 1868) Garoupa zebrada Skeweband-klipkabeljou Daghma Hamour Lapu-lapu Pugapo Grouper Obliquebanded grouper Oblique-banded grouper Oblique-banded rockcod Radiant rockcod Mérou zébré Lokos Kakehashihata Kerapu Kerapu jejari serong Manjel kaleva Lapu lapu Mero acebrado Bulang Cabot de fond Gerape rimung Geurape rimueng Inkada Kerapu harimau Kurapu Raja laveya Yau-paan 云纹石斑鱼 电纹石斑鱼 石斑 过鱼 雲紋石斑魚 電紋石斑魚 鲙 Epinephelus döderleinii Franz, 1910 Epinephelus doederleinii Franz, 1910 Epinephelus morrhua (non Valenciennes, 1833) Epinephelus radians (Day, 1868) Epinephelus radiates (Day, 1868) Epinephilus radiatus (Day, 1868) Mycteroperca radiatus (Day, 1868) Serranus brunneus (non Bloch, 1793) Serranus morrhua (non Valenciennes, 1833) Serranus radiatus Day, 1868 1993-02-12 2014-11-26 Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter0 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) Eschmeyer, W.N.0 Catalog of the genera of recent fishes. California Academy of Sciences, San Francisco, USA. 697 p. (Ref. 1830) Allen, G.R. and R. Swainston0 The marine fishes of north-western Australia: a field guide for anglers and divers. Western Australian Museum, Perth. 201 p. (Ref. 3132) Heemstra, P.C. and J.E. Randall0 Serranidae. In W. Fischer and G. Bianchi (eds.). FAO species identification sheets for fishery purposes. Western Indian Ocean (fishing area 51). Rome, FAO. Vol. IV. 241 p. (Ref. 3153) Heemstra, P.C. and J.E. Randall0 Serranidae. p. 509-537. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 4319) Randall, J.E. and P.C. Heemstra0 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787) Fischer, W., I. Sousa, C. Silva, A. de Freitas, J.M. Poutiers, W. Schneider, T.C. Borges, J.P. Feral and A. Massinga0 Fichas FAO de identificaçao de espécies para actividades de pesca. Guia de campo das espécies comerciais marinhas e de águas salobras de Moçambique. Publicaçao preparada em collaboraçao com o Instituto de Investigaçao Pesquiera de Moçambique, com financiamento do Projecto PNUD/FAO MOZ/86/030 e de NORAD. Roma, FAO. 1990. 424 p. (Ref. 5213) Heemstra, P.C. and J.E. Randall0 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) Gloerfelt-Tarp, T. and P.J. Kailola0 Trawled fishes of southern Indonesia and northwestern Australia. Australian Development Assistance Bureau, Australia, Directorate General of Fishes, Indonesia, and German Agency for Technical Cooperation, Federal Republic of Germany. 407 p. (Ref. 5978) Kailola, P.J.0 The fishes of Papua New Guinea: a revised and annotated checklist. Vol. II Scorpaenidae to Callionymidae. Research Bulletin No. 41, Research Section, Dept. of Fisheries and Marine Resources, Papua New Guinea. (Ref. 6192) Fouda, M.M. and G.V. Hermosa Jr.0 A checklist of Oman fishes. Sultan Qaboos University Press, Sultanate of Oman. 42 p. (Ref. 6365) Katayama, M.0 Family Serranidae. p. 126-138. In H. Masuda, K. Amaoka, C. Araga, T. Uyeno, and T. Yoshino (eds). The fishes of the Japanese archipelago. Second Edition. Tokai University Press, Tokyo. 456 p. (Ref. 6854) Paxton, J.R., D.F. Hoese, G.R. Allen and J.E. Hanley0 Pisces. Petromyzontidae to Carangidae. Zoological Catalogue of Australia, Vol. 7. 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(Ref. 31667) Fricke, R.0 Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez): an annotated checklist, with descriptions of new species. Koeltz Scientific Books, Koenigstein, Theses Zoologicae, Vol. 31:759 p. (Ref. 33390) Randall, J.E. and K.K.P. Lim (eds.)0 A checklist of the fishes of the South China Sea. Raffles Bull. Zool. Suppl. (8):569-667. (Ref. 36648) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Heemstra, P.C. and J.E. Randall0 Serranidae. p. 2442-2547. In K.E. Carpenter and V.H. Niem (eds.) FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific. Vol. 4. Bony fishes part 2 (Mugilidae to Carangidae). Rome, FAO. 2069-2790 p. (Ref. 39231) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Anonymous0 Fish collection of the University of the Philippines in the Visayas Museum. UPV Museum. (Ref. 43309) Kapoor, D., R. Dayal and A.G. Ponniah0 Fish biodiversity of India. National Bureau of Fish Genetic Resources Lucknow, India.775 p. (Ref. 45255) Huang, Z.0 Marine species and their distribution in China's seas. p. 404-463. Vertebrata. Smithsonian Institution, Florida, USA. 598 p. (Ref. 47843) Hardy, J.D. Jr. (comp.)0 Coral reef fish species. NOAANational Oceanographic Data Center. NODC Coral Reef Data and Information Management System. USA. 537 p. (Ref. 48497) Khalaf, M.A. and A.M. Disi0 Fishes of the Gulf of Aqaba. Marine Science Station, Aqaba, Jordan. 252 p. (Ref. 48643) Letourneur, Y., P. Chabanet, P. Durville, M. Taquet, E. Teissier, M. Parmentier, J.-C. Quéro and K. Pothin0 An updated checklist of the marine fish fauna of Reunion Island, south-western Indian Ocean. Cybium 28(3):199-216. (Ref. 53568) Chen, C.-H.0 Checklist of the fishes of Penghu. 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Zoological Survey of India, Andaman and Nicobar Regional Centre, Haddo, Port Blair. (Ref. 87596) Craig, M.T., YJ. Sadovy de Mitcheson and P.C. Heemstra0 Groupers of the world: a field and market guide. North America: CRC Press/Taylor and Francis Group, xix, 356 p., A47 pages appendix. (Ref. 89707) Taquet, M. and A. Diringer0 Mérous de l'Océan Indien. La Réunion: Azalées Editions. 103 p. (Ref. 89967) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632) Stobbs, R.E.0 Comorian fish names with a preliminary list of Malagasy common names. Investigational Report No. 35 (Ref. 97705) Kandula, S., K.V.L. Shrikanya and V.A. Iswarya Deepti0 Species diversity and some aspects of reproductive biology and life history of groupers (Pisces: Serranidae: Epinephelinae) off the central eastern coast of India. Mar. Biol. Res. 11(1):18-33. (Ref. 97769) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257) Sujatha, K., K.V.L. Shrikanya and P. Padmavathi0 Length-weight relationship of four species of <i>Epinephelus</i> Bloch, 1793 in the catches off Visakhapatnam, east coast of India. J. Mar. Biol. Ass. India 52(1):96-98. (Ref. 116276) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272) Ma, K.Y. and M.T. Craig0 An inconvenient monophyly: an update on the taxonomy of the groupers (Epinephelidae). Copeia 106(3):443-456. (Ref. 119507) IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435) Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577) FB-pic-7360-7339 http://purl.org/dc/dcmitype/StillImage image/gif FAO 1996-01-15 1996-09-10 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://www.fishbase.us/images/species/Eprad_u0.gif http://www.fishbase.us/images/thumbnails/gif/tn_Eprad_u0.gif FB-pic-7360-9609 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-03 2000-11-09 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology http://www.fishbase.us/images/species/Eprad_u2.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Eprad_u2.jpg FB-pic-7360-9610 http://purl.org/dc/dcmitype/StillImage image/jpeg John E. Randall 1996-03-03 2010-07-22 http://creativecommons.org/licenses/by-nc/3.0/ This work is licensed under a Creative Commons Attribution-Noncommercial 3.0 Unported License. Image can be freely copied and altered but may be used only for non-commercial purposes. Original author and source must be properly acknowledged WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution market; 1.5 lbs. Locality: Naha, Okinawa, Ryukyu Is.. http://www.fishbase.us/images/species/Eprad_u3.jpg http://www.fishbase.us/images/thumbnails/jpg/tn_Eprad_u3.jpg Locality: Naha, Okinawa, Ryukyu Is.. FB-GeneralDescr-7360 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2014-11-26 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 Adults are found at depths between 80 to 383 m; juveniles in 18 to 20 m (Ref. 5222), over hard substrates (Ref. 89707). The species is apparently rare (Ref. 5222). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-DiagnosticDescr-7360 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-13 00:00:00 2013-10-21 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size Distinguished by the following characteristics: tan body color with 5 oblique dark-edged pale bands; dark bands with scattered small black spots and pale blotches, especially dorsally; pale interspaces with small dark brown spots, mainly arranged in series along the middle of the interspaces; adults with dark brown line from the lower edge of the eye to the edge of the subopercle; faint dark band along the maxillary groove and continuing to the edge of the interopercle; dorsal fin and caudal fin covered with small dark spots; juveniles dark brown with black-edged pale brown bands enclosing numerous small black spots; body depth contained 2.6-3.0 times in SL; head length 2.1-2.3 times in SL; nearly flat interorbital, dorsal head profile slightly convex; 2-5 distinctly enlarged serrae on preopercle angle; upper edge of operculum almost straight; adults posterior nostril diameter 2-3 times that of anterior nostrils; maxilla reaches to or past vertical at rear edge of eye; 2 rows of teeth on midlateral part of lower jaw (Ref. 89707). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Morphology-7360 http://purl.org/dc/dcmitype/Text text/html Rodolfo B. Reyes 1994-06-13 00:00:00 2013-10-21 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Dorsal spines (total): 11; Dorsal soft rays (total): 13 - 15; Analspines: 3; Analsoft rays: 8 Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Distribution-7360 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2025-11-06 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Indo-West Pacific: spotty distribution from the Red Sea to Japan and Papua New Guinea. Referred to as Epinephelus morrhua morrhua in Japan (Ref. 5978). Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222) FB-Uses-7360 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2014-11-26 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution fisheries: minor commercial; price category: very high; price reliability: questionable: based on ex-vessel price for species in this genus Coppola, S.R., W. Fischer, L. Garibaldi, N. Scialabba and K.E. Carpenter 1994 SPECIESDAB: Global species database for fishery purposes. User's manual. FAO Computerized Information Series (Fisheries). No. 9. Rome, FAO. 103 p. (Ref. 171) FB-TrophicStrategy-7360 http://purl.org/dc/dcmitype/Text text/html Drina Sta. Iglesia 2001-06-07 00:00:00 2024-02-06 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ radiatus Found on the continental shelf (Ref. 75154). Adults are found at depths between 80 to 383 m; juveniles in 18 to 20 m (Ref. 5222). Attain 70 cm TL (Ref. 58472). The species is apparently rare (Ref. 5222). Dwells in rocky substrates. Juveniles prefer shallow waters of about 18-20m deep. Reportedly enters estuaries despite intolerance of low salinities. Feeds on small fishes, crustaceans, crabs and lobsters (Ref. 9773). Masuda, H. and G.R. Allen 1993 Meeresfische der Welt - Groß-Indopazifische Region. Tetra Verlag, Herrenteich, Melle. 528 p. (Ref. 9137) FB-Size-7360 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2014-11-26 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size 70.0 cm SL (male/unsexed; (Ref. 4787)) Randall, J.E. and P.C. Heemstra 1991 Revision of Indo-Pacific groupers (Perciformes: Serranidae: Epinephelinae), with descriptions of five new species. Indo-Pac. Fish. (20):332 p. (Ref. 4787) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-7360 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 1993-02-12 00:00:00 2014-11-26 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=7360 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat demersal; marine; depth range 18 - 383 m (Ref. 5222) Heemstra, P.C. and J.E. Randall 1993 FAO Species Catalogue. Vol. 16. Groupers of the world (family Serranidae, subfamily Epinephelinae). An annotated and illustrated catalogue of the grouper, rockcod, hind, coral grouper and lyretail species known to date. Rome: FAO. FAO Fish. Synop. 125(16):382 p. (Ref. 5222)