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	<dc:identifier>FB-7960</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7960</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rajiformes</dwc:Order>
	<dwc:Family>Rajidae</dwc:Family>
	<dwc:Genus>Dipturus</dwc:Genus>
	<dwc:ScientificName>Dipturus pullopunctatus (Smith, 1964)</dwc:ScientificName>
	<commonName xml:lang='af'>Rog</commonName>
	<commonName xml:lang='en'>Skate</commonName>
	<commonName xml:lang='en'>Slime skate</commonName>
	<commonName xml:lang='et'>Libe tiibrai</commonName>
	<commonName xml:lang='fr'>Raie baveuse</commonName>
	<commonName xml:lang='es'>Raya limosa</commonName>
	<commonName xml:lang=''>Slijmrog</commonName>
	<commonName xml:lang=''>暗斑鰩</commonName>
	<commonName xml:lang=''>暗斑鳐</commonName>
	<synonym relationship='synonym'>Dipturus pullopunctata (Smith, 1964)</synonym>
	<synonym relationship='misapplied name'>Raia batis (non Linnaeus, 1758)</synonym>
	<synonym relationship='synonym'>Raia pullopunctata Smith, 1964</synonym>
	<synonym relationship='misapplied name'>Raja campbelli (non Wallace, 1967)</synonym>
	<synonym relationship='synonym'>Raja pullopunctata Smith, 1964</synonym>
	<dcterms:created>1993-04-21</dcterms:created>
	<dcterms:modified>2018-11-20</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=7960'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2708&amp;speccode=7960'>Hulley, P.A.0 Rajidae. p. 115-127. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 2708)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=7960'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7458&amp;speccode=7960'>Ebert, D.A., P.D. Cowley and L.J.V. Compagno0 A preliminary investigation of the feeding ecology of skates (Batoidea: Rajidae) off the west coast of southern Africa. S. Afr. J. Mar. Sci. 10:71-81. (Ref. 7458)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10311&amp;speccode=7960'>Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman &amp; Hall, London. (Ref. 10311)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11228&amp;speccode=7960'>Heemstra, P.C.0 Additions and corrections for the 1995 impression. p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin. (Ref. 11228)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11284&amp;speccode=7960'>Stehmann, M.0 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=7960'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27121&amp;speccode=7960'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265p. (Ref. 27121)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=7960'>McEachran, J.D. and K.A. Dunn0 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=7960'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=7960'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=7960'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='92-5-104345-0' url='http://www.fishbase.org/references/FBRefSummary.php?id=36731&amp;speccode=7960'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265 p. (Ref. 36731)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=40919&amp;speccode=7960'>Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=7960'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=46206&amp;speccode=7960'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of March 2003. Database dump. Catalog databases as made available to FishBase in March 2003. (Ref. 46206)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=49896&amp;speccode=7960'>Sulikowski, J.A., M.D. Morin, S.H. Suk and W.H. Howell0 Age and growth estimates of the winter skate (&lt;i&gt;Leucoraja ocellatal&lt;/i&gt;) in the western Gulf of Maine. Fish Bull. 101(2):405-413. (Ref. 49896)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=7960'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52207&amp;speccode=7960'>Wamsley-Hart, S.A., W.H.H. Sauer and C.D. Buxton0 The biology of the skates &lt;i&gt;Raja wallacei&lt;/i&gt; and &lt;i&gt;R. pullopunctata&lt;/i&gt; (Batoidea: Rajidae) on the Agulhas Bank, South Africa. S. Afr. J. Mar. Sci. 21:165-179. (Ref. 52207)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53748&amp;speccode=7960'>Sulikowski, J.A., J. Kneebone, S. Elzey, J. Jurek, P.D. Danley, W. Huntting Howell and P.C.W. Tsang0 Age and growth estimates of the thorny skate (&lt;i&gt;Amblyraja radiata&lt;/i&gt;) in the western Gulf of Maine. Fish. Bull. 103:161-168. (Ref. 53748)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=7960'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80491&amp;speccode=7960'>Ebert, D.A., L.J.V. Compagno and P.D. Cowley0 Aspects of the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern Africa. ICES Journal of Marine Science 65(1):81-102. (Ref. 80491)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=84081&amp;speccode=7960'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated internet version of 06 May 2010. Catalog databases of CAS cited in FishBase (website). (Ref. 84081)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=7960'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=7960'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor0 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=7960'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1993-04-21 00:00:00</dcterms:created>
	<dcterms:modified>2018-11-20 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7960</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Found on soft bottoms of the outer shelf and upper slope (Ref. 5578).  Feeds on bony fish, crabs, shrimp, mysids, mantis shrimp, euphausiids, bivalves and cuttlefish (Ref. 5578).  Oviparous (Ref. 50449). Eggs have horn-like projections on the shell (Ref. 205). Males reaches maturity at 88-96 cm TL, females possibly slightly larger off Western than Southern coast of South Africa; hatching size at ca. 19 cm TL (Ref. 114953).  Commonly taken by hake trawlers (Ref. 5578).  Extremely slippery when caught (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=7960'>McEachran, J.D. and K.A. Dunn 1998 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-11-25 00:00:00</dcterms:created>
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	<dc:description>A striking two-toned longnose skate, medium brown above and abruptly dark grey below, with a large, conspicuous, dark brown blotch on the upper base of each pectoral fin (Ref. 5578).  Snout moderately elongated and bluntly triangular, and tail stout, not conspicuously swollen and about equal to body length; pectoral disc with broadly rounded corners and smooth underside without denticles; thorns at nape and along midline of back and tail to 1st dorsal fin, nape spine enormous in young (Ref. 5578).  Hatchlings and juveniles with wide-spaced black spots on upper disc, inconspicuous or lost in adolescents and adults; adults sometimes with numerous small white spots (Ref. 5578).  Underside with numerous black pores (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=7960'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dc:description>Southeast Atlantic:  known only from Lüderitz, Namibia to Port Alfred, Eastern Cape (South Africa)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=7960'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<dc:title>Importance</dc:title>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2708&amp;speccode=7960'>Hulley, P.A. 1986 Rajidae. p. 115-127. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 2708)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=7960&amp;StockCode=8271</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Migration</subject>
	<dc:description>Oviparous, paired eggs are laid.  Embryos feed solely on yolk (Ref. 50449).  Eggs have horn-like projections on the shell (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=7960'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>130 cm TL (male/unsexed; (Ref. 114953))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=114953&amp;speccode=7960'>Last, P.R., W.T. White, M.R. de Carvalho, B. Séret, M.F.W. Stehmann and G.J.P. Naylor 2016 Rays of the world. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. (Ref. 114953)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-7960</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1993-04-21 00:00:00</dcterms:created>
	<dcterms:modified>2018-11-20 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=7960</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>demersal; marine; depth range 50 - 457 m (Ref. 5578)</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=7960'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject>
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