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	<dc:identifier>FB-8361</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Copadichromis</dwc:Genus>
	<dwc:ScientificName>Copadichromis mbenjii Konings, 1990</dwc:ScientificName>
	<commonName xml:lang='en'>Happy</commonName>
	<commonName xml:lang='fi'>Keltakylkiutaka</commonName>
	<commonName xml:lang=''>Utaka</commonName>
	<commonName xml:lang=''>本氏桨鳍丽鱼</commonName>
	<commonName xml:lang=''>本氏槳鰭麗魚</commonName>
	<synonym relationship='synonym'>Copadichromis mbenji Konings, 1990</synonym>
	<synonym relationship='misapplied name'>Haplochromis quadrimaculatus (non Regan, 1922)</synonym>
	<dcterms:created>1993-08-31</dcterms:created>
	<dcterms:modified>2008-10-31</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4979&amp;speccode=8361'>Maréchal, C.0 Copadichromis. p. 51-58. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 4979)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5595&amp;speccode=8361'>Konings, A.0 Ad Konings's book of cichlids and all the other fishes of Lake Malawi. T.F.H. Publications, Inc. 495 p. (Ref. 5595)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12468&amp;speccode=8361'>Baensch, H.A. and R. Riehl0 Aquarien Atlas. Band 4. Mergus Verlag GmbH, Verlag für Natur-und Heimtierkunde, Melle, Germany. 864 p. (Ref. 12468)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8361'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8361'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=33486&amp;speccode=8361'>Nsiku, E.0 Changes in the fisheries of Lake Malawi, 1976-1996: ecosystem-based analysis. University of British Columbia, Vancouver, Canada. 217 p. M.Sc. thesis. (Ref. 33486)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=8361'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55463&amp;speccode=8361'>Snoeks, J. (ed.)0 The cichlid diversity of Lake Malawi/Nyasa/Niassa: identification, distribution and taxonomy. Cichlid Press, El Paso, USA, 360p. (Ref. 55463)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55953&amp;speccode=8361'>Chisambo, J. and J. Snoeks0 Key to the deep-bodied, spotted &lt;i&gt;Copadichromis&lt;/i&gt; species. p.261-265. In Snoeks, J. (ed.) The cichlid diversity of Lake Malawi/Nyasa/Niassa: identification, distribution and taxonomy. Cichlid Press, El Paso, USA, 360p. (Ref. 55953)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56787&amp;speccode=8361'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Updated database version of April 2006. Database dump. Catalog databases as made available to FishBase in April 2006. (Ref. 56787)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57549&amp;speccode=8361'>Stauffer, J.R. and A.F. Konings0 Review of &lt;i&gt;Copadichromis&lt;/i&gt; (Teleostei: Cichlidae) with the description of a new genus and six new species. Ichthyol. Explor. Freshwat. 17(1):9-42. (Ref. 57549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8361'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=8361'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-8361</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1993-08-31 00:00:00</dcterms:created>
	<dcterms:modified>2008-10-31 21:46:09</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<dc:description>Occurs at the sand/rock interface (Ref. 57549). Found at an average depth of 15 m (Ref. 5595). Feeds on plankton (Ref. 5595, 57549) and benthic invertebrates (Ref. 57549). Territorial males build a bowl-shaped bower beneath and/or adjacent to a rock, 18-42cm in diameter and a 4-7 cm high rim; females occur in schools numbering up to 50 individuals, remaining within 2m above the bottom (Ref. 57549).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4979&amp;speccode=8361'>Maréchal, C. 1991 Copadichromis. p. 51-58. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 4979)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-8361</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2008-10-30 20:40:40</dcterms:created>
	<dcterms:modified>2008-10-30 20:40:40</dcterms:modified>
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	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Diagnosis: 3 lateral spots on the body; supra-pectoral spot 2-2.5 times diameter of supra-anal spot; 12-15 gill rakers on the first ceratobranchial; breeding males with yellow breast, wide yellow marginal band in anal fin and a distinct yellow dorsal fin margin (Ref. 57549).Description: dorsal head profile convex, with a slight concavity between eye and snout tip in large males, convex in females and immatures; gape inclination of mouth about 40° to horizontal; jaws isognathous; teeth in 3 rows on dentary and premaxilla, with outer row teeth bicuspid in females and immatures and unequally bicuspid tending to unicuspid in mature males, while inner rows are unicuspid; lower pharyngeal jaw small, all teeth bicuspid, posterior 2 central teeth sometimes slightly enlarged; gill-rakers on ceratobranchial short and broad-based; 3 rows of teeth in upper and lower jaw; 2-3 scales rows on cheek; 1-3 pored scales posterior to lateral line (Ref. 57549).Coloration: breeding males: head blue; throat white with gray patches; lateral portion of body light-blue suffused with orange, anterior portion of scales orange/brown and posterior edge of scales light-blue; nape orange/brown; breast yellow with blue/gray spots; belly gray with blue/white highlights; dorsal membranes blue with orange/brown markings, membranes between rays with orange/brown spots and streaks; dorsal spines and rays blue proximally and gray/blue distally; dorsal fin with narrow black submarginal band with white lappets and yellow tips; caudal fin with blue rays and yellow/orange membranes; proximal half of anal fin blue/black, distal half yellow; posterior half of anal fin with white ocelli; pectoral fin clear; pelvic fins yellow/brown with white leading edge and dark band between 1st and 2nd ray (Ref. 57549). Females: head gray/purple; cheeks silvery with purple highlights; opercle silver with brown and purple highlights and black opercular spot; throat white; lateral portion of body with 3 black spots: supra-pectoral spot below 9-lOth scale in contact with upper lateral line and about 2 times size of supra-anal spot; supra-anal spot at 21-23nd scale above and in contact with lower lateral line and covering 2-3 scales; caudal spot same diameter as supra-anal spot on caudal base; above upper lateral line bronze ground color with purple highlights, below upper lateral line silver ground color with center of each scale yellow/brown; 4 vertical bars between supra-pectoral spot and dorsal-fin origin; breast and belly white; dorsal fin clear with orange/brown spots, white lappets, and orange/brown tips; caudal fin gray/yellow with orange/brown spots; proximal 2/3 of anal fin light gray, distal 1/3 clear with orange/brown ocelli and narrow orange/brown edge; pectoral fin clear; pelvic fin clear with gray membrane between first 2 rays (Ref. 57549).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57549&amp;speccode=8361'>Stauffer, J.R. and A.F. Konings 2006 Review of &lt;i&gt;Copadichromis&lt;/i&gt; (Teleostei: Cichlidae) with the description of a new genus and six new species. Ichthyol. Explor. Freshwat. 17(1):9-42. (Ref. 57549)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-8361</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2008-10-30 20:40:40</dcterms:created>
	<dcterms:modified>2008-10-30 20:40:40</dcterms:modified>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<dc:description>Dorsal spines (total): 16 - 18; Dorsal soft rays (total): 10 - 11;  Analspines: 3; Analsoft rays: 8 - 9</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57549&amp;speccode=8361'>Stauffer, J.R. and A.F. Konings 2006 Review of &lt;i&gt;Copadichromis&lt;/i&gt; (Teleostei: Cichlidae) with the description of a new genus and six new species. Ichthyol. Explor. Freshwat. 17(1):9-42. (Ref. 57549)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-8361</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1993-08-31 00:00:00</dcterms:created>
	<dcterms:modified>2019-02-04 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Africa:  Endemic to Mbenji Island, Lake Malawi.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4979&amp;speccode=8361'>Maréchal, C. 1991 Copadichromis. p. 51-58. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 4979)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-8361</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1993-08-31 00:00:00</dcterms:created>
	<dcterms:modified>2008-10-31 21:46:09</dcterms:modified>
	<dc:title>Importance</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>fisheries: commercial; aquarium: commercial</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5595&amp;speccode=8361'>Konings, A. 1990 Ad Konings's book of cichlids and all the other fishes of Lake Malawi. T.F.H. Publications, Inc. 495 p. (Ref. 5595)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5595&amp;speccode=8361'>Konings, A. 1990 Ad Konings's book of cichlids and all the other fishes of Lake Malawi. T.F.H. Publications, Inc. 495 p. (Ref. 5595)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-8361</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1993-08-31 00:00:00</dcterms:created>
	<dcterms:modified>2008-10-31 21:46:09</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>14.0 cm TL (male/unsexed; (Ref. 4979))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4979&amp;speccode=8361'>Maréchal, C. 1991 Copadichromis. p. 51-58. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 4979)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-8361</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1993-08-31 00:00:00</dcterms:created>
	<dcterms:modified>2008-10-31 21:46:09</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8361</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>benthopelagic; freshwater; pH range: 7.3 - 8.4; dH range: 5 - 30; depth range 7 - 19 m (Ref. 57549)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57549&amp;speccode=8361'>Stauffer, J.R. and A.F. Konings 2006 Review of &lt;i&gt;Copadichromis&lt;/i&gt; (Teleostei: Cichlidae) with the description of a new genus and six new species. Ichthyol. Explor. Freshwat. 17(1):9-42. (Ref. 57549)</reference></dataObject>
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