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	<dc:identifier>FB-8921</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Coptodon</dwc:Genus>
	<dwc:ScientificName>Coptodon louka (Thys van den Audenaerde, 1969)</dwc:ScientificName>
	<commonName xml:lang='fi'>Keltatilapia</commonName>
	<commonName xml:lang=''>卢卡非鲫</commonName>
	<commonName xml:lang=''>盧卡非鯽</commonName>
	<synonym relationship='synonym'>Tilapia louka Thys van den Audenaerde, 1969</synonym>
	<synonym relationship='misapplied name'>Tilapia melanopleura (non Duméril, 1861)</synonym>
	<synonym relationship='misapplied name'>Tilapia rheophila (non Daget, 1962)</synonym>
	<dcterms:created>1994-03-04</dcterms:created>
	<dcterms:modified>2024-03-22</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=8921'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7378&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 714-779. In C. Levêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest. Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, 902 p. (Ref. 7378)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=9932&amp;speccode=8921'>Pouyaud, L. and J.-F. Agnèse0 Phylogenetic relationships between 21 species of three tilapiine genera &lt;i&gt;Tilapia&lt;/i&gt;, &lt;i&gt;Sarotherodon&lt;/i&gt; and &lt;i&gt;Oreochromis&lt;/i&gt; using allozyme data. J. Fish Biol. 47(1):26-38. (Ref. 9932)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12468&amp;speccode=8921'>Baensch, H.A. and R. Riehl0 Aquarien Atlas. Band 4. Mergus Verlag GmbH, Verlag für Natur-und Heimtierkunde, Melle, Germany. 864 p. (Ref. 12468)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=8921'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13331&amp;speccode=8921'>Paugy, D., K. Traoré and P.S. Diouf0 Faune ichtyologique des eaux douces d'Afrique de l'Ouest. p.35-66. In G.G. Teugels, J.F. Guégan and J.J. Albaret (eds.) Biological diversity of African fresh-and brackish water fishes. Geographical overviews presented at the PARADI Symposium, Senegal, 15-20 November 1993. Ann. Mus. R. Afr. Centr., Sci. Zool., 275:177 p. (Ref. 13331)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8921'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8921'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=8921'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36297&amp;speccode=8921'>Hugueny, B. and M. Pouilly0 Morphological correlates of diet in an assemblage of West African freshwater fishes. J. Fish Biol. 54(6):1310-1325. (Ref. 36297)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=8921'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=8921'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=8921'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57398&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 520-600. In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturelle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57398)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8921'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=8921'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=8921'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8921'>Dunz, A.R. and U.K. Schliewen0 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=8921'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=8921'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=8921'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-8921</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-22 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Seems to prefer brackish regions and coastal lagoons; probably has a high tolerance for elevated salinity levels; herbivorous; pair-bonding, open substrate spawner, with both parents guarding the brood (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93285&amp;speccode=8921'>Dunz, A.R. and U.K. Schliewen 2013 Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as "&lt;i&gt;Tilapia&lt;/i&gt;". Mol. Phylogenet. Evol. 68(1):64-80. (Ref. 93285)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-8921</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2008-11-21 22:02:48</dcterms:created>
	<dcterms:modified>2018-01-09 20:46:34</dcterms:modified>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<dc:description>Diagnosis: lower pharyngeal bone about as long as broad, and with anterior lamella shorter than toothed area; median pharyngeal teeth not broadened; dorsal fin with 13-16 spines and 12-14 soft rays; soft dorsal and upper tip of caudal bordered with orange-red; "tilapia" spot clearly marked even in adults; dorsal and caudal fins not or feebly blotched (Ref. 53405).Description: deep-bodied; mouth small; caudal fin slightly rounded (Ref. 52307). Outer jaw teeth bicuspid and non-spatulate, inner jaw teeth tricuspid, posterior pharyngeal teeth tricuspid with the lower cusp rather small; micro-gillrakers present; scales cycloid; pectoral region covered with very small scales (Ref. 53405).Coloration: in life: adults: ground colour greyish to silvery, more whitish on belly and rather dark grey on back, occiput and snout; sides with 5-6 faint cross bars and horizontal band; lower part of cheeks and subopercle bright orange-red, and traces of this colouration on opercle and breast; first 1/4 of spinous part of dorsal fin uniformly grey, posterior part with 2-3 rows of light oblique bars between spines; soft part of dorsal fin with irregular light spots; well visible "tilapian" spot; distal margin of dorsal fin black on spinous part, orange-red on soft part; anal fin dark grey, with distal margin lighter and some light spots posteriorly; pelvic fins with black anterior margin: caudal fin with orange-red upper angle and light spots forming a network on proximal part and transverse series on distal part (Ref. 53405). Dorsal, caudal and anal fins often with red margin, sometimes their bases also appear reddish (Ref. 52307). Juveniles: less colourful than adults; anterior part of spinous dorsal fin and distal areas of soft dorsal and caudal fins darker, with much more black between branched rays (Ref. 53405).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-8921</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
	<dcterms:created>2008-11-21 22:02:48</dcterms:created>
	<dcterms:modified>2018-01-09 20:46:34</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Dorsal spines (total): 13 - 16; Dorsal soft rays (total): 12 - 14;  Analspines: 3; Analsoft rays: 8 - 10</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-8921</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
	<dcterms:modified>2019-02-04 00:00:00</dcterms:modified>
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	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Africa:  Guinea Bissau (Corubal River) to Liberia (Lofa River)(Ref. 52307, 53405).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
	<dc:identifier>FB-Size-8921</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-22 00:00:00</dcterms:modified>
	<dc:title>Max. size</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>25.0 cm TL (male/unsexed; (Ref. 5163))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5163&amp;speccode=8921'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 1991 Tilapia. p. 482-508. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5163)</reference></dataObject><dataObject>
	<dc:identifier>FB-Threats-2</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
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	<dc:rights></dc:rights>
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	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
	<mediaURL></mediaURL>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-8921</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-22 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8921</dc:source>
	<dc:description>benthopelagic; freshwater; brackish</dc:description>
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