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	<dc:identifier>FB-8938</dc:identifier>
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	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rajiformes</dwc:Order>
	<dwc:Family>Rajidae</dwc:Family>
	<dwc:Genus>Rajella</dwc:Genus>
	<dwc:ScientificName>Rajella leopardus (von Bonde &amp; Swart, 1923)</dwc:ScientificName>
	<commonName xml:lang='af'>Rog</commonName>
	<commonName xml:lang='en'>Leopard skate</commonName>
	<commonName xml:lang='en'>Skate</commonName>
	<commonName xml:lang='et'>Leopard-väikerai</commonName>
	<commonName xml:lang='fr'>Raie léopard</commonName>
	<commonName xml:lang='es'>Raya leopardo</commonName>
	<commonName xml:lang=''>Luipaardrog</commonName>
	<commonName xml:lang=''>豹紋鰩</commonName>
	<commonName xml:lang=''>豹纹鳐</commonName>
	<synonym relationship='synonym'>Raia leopardus von Bonde &amp; Swart, 1923</synonym>
	<synonym relationship='misapplied name'>Raja barnardi (non Norman, 1935)</synonym>
	<synonym relationship='synonym'>Raja leopardus von Bonde &amp; Swart, 1923</synonym>
	<synonym relationship='synonym'>Rajella leoparda (von Bonde &amp; Swart, 1923)</synonym>
	<dcterms:created>1994-03-04</dcterms:created>
	<dcterms:modified>2021-10-15</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=8938'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2708&amp;speccode=8938'>Hulley, P.A.0 Rajidae. p. 115-127. In M.M. Smith and P.C. Heemstra (eds.) Smiths' sea fishes. Springer-Verlag, Berlin. (Ref. 2708)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4426&amp;speccode=8938'>Stehmann, M.0 Rajidae. p. 29-50. In J.C. Quero, J.C. Hureau, C. Karrer, A. Post and L. Saldanha (eds.) Check-list of the fishes of the eastern tropical Atlantic. Junta Nacional de Investigaçao Cientifica e Tecnológica, Lisbon, Portugal. Vol. 1. (Ref. 4426)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=8938'>Compagno, L.J.V., D.A. Ebert and M.J. Smale0 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7458&amp;speccode=8938'>Ebert, D.A., P.D. Cowley and L.J.V. Compagno0 A preliminary investigation of the feeding ecology of skates (Batoidea: Rajidae) off the west coast of southern Africa. S. Afr. J. Mar. Sci. 10:71-81. (Ref. 7458)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=10311&amp;speccode=8938'>Fritzsch, B. and P. Moller0 A history of electroreception. p. 39-55. In P. Moller (ed.) Electric fishes:history and behavior. Fish and Fisheries Series 17. Chapman &amp; Hall, London. (Ref. 10311)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11228&amp;speccode=8938'>Heemstra, P.C.0 Additions and corrections for the 1995 impression. p. v-xv. In M.M. Smith and P.C. Heemstra (eds.) Revised Edition of Smiths' Sea Fishes. Springer-Verlag, Berlin. (Ref. 11228)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=11284&amp;speccode=8938'>Stehmann, M.0 First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Arch. Fish. Mar. Res. 43(1):1-119. (Ref. 11284)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=26282&amp;speccode=8938'>Eschmeyer, W.N. (ed.)0 Catalog of fishes. Special Publication, California Academy of Sciences, San Francisco. 3 vols. 2905 p. (Ref. 26282)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27121&amp;speccode=8938'>Bianchi, G., K.E. Carpenter, J.-P. Roux, F.J. Molloy, D. Boyer and H.J. Boyer0 FAO species identification guide for fishery purposes. Field guide to the living marine resources of Namibia. FAO, Rome. 265p. (Ref. 27121)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=8938'>McEachran, J.D. and K.A. Dunn0 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=8938'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=8938'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=8938'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36670&amp;speccode=8938'>Anonymous0 Fish collection database of the J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. J.L.B. Smith Institute of Ichthyology, Grahamstown, South Africa. (Ref. 36670)</reference>
	<reference isbn='84-931926-0-0.' url='http://www.fishbase.org/references/FBRefSummary.php?id=39531&amp;speccode=8938'>Crespo, J., J. Gajate and R. Ponce0 Clasificación científica e identificación de nombres vernáculos existentes en la base de datos de seguimiento informático de recursos naturales oceánicos. Instituto Español de Oceanografia, Madrid, Spain. 414 p. (Ref. 39531)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41249&amp;speccode=8938'>Ishihara, H.0 Study on the systematics and fishery resources of the North Pacific skates (Pisces: Chondrichthyes: Rajidae). Tokyo University. 186 p. Ph.D. dissertation. (Ref. 41249)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41250&amp;speccode=8938'>Bor, P.0 Egg-capsules of sharks and skates. www.rajidae.tmfweb.nl/rogtabel.html (Ref. 41250)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=8938'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=8938'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=8938'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58748&amp;speccode=8938'>Macpherson, E.0 Influence of geographical distribution, body size and diet on population density of benthic fishes off Namibia (South West Africa). Mar. Ecol. Prog. Ser. 50:295-299. (Ref. 58748)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=75549&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=80491&amp;speccode=8938'>Ebert, D.A., L.J.V. Compagno and P.D. Cowley0 Aspects of the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern Africa. ICES Journal of Marine Science 65(1):81-102. (Ref. 80491)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96993&amp;speccode=8938'>Weigmann, S., M.F.W. Stehmann and R. Thiel0 &lt;i&gt;Rajella paucispinosa&lt;/i&gt; n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the western Indian Ocean off South Mozambique, and a revised generic diagnosis. Zootaxa 3847(3):359-387. (Ref. 96993)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=8938'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=8938'>Weigmann, S.0 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=115257&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118272&amp;speccode=8938'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118749&amp;speccode=8938'>Last, P.R., S. Weigmann and L. Yang0 Changes to the nomenclature of the skates (Chondrichthyes: Rajiformes). In: Last and Yearsley (eds.). Rays of the World: Supplementary Information. CSIRO Special Publication. 11-34. (Ref. 118749)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=130435&amp;speccode=8938'>IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dc:description>Found in deeper shelf and slope waters (Ref. 4426).  Feed on bottom crustaceans, dragonets, rattails, hake, lanternfish, sea pens, cuttlefish and polychaete worms (Ref. 5578).  Oviparous.  Distinct pairing with embrace.  Young may tend to follow large objects, such as their mother (Ref. 205).  Eggs are oblong capsules with stiff pointed horns at the corners deposited in sandy or muddy flats (Ref. 205).  Egg capsules are 7.4 cm long and 7.4 cm wide (Ref. 41249).  Caught by hake trawlers (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=8938'>McEachran, J.D. and K.A. Dunn 1998 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-11-24 00:00:00</dcterms:created>
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	<dc:description>Large skate with a moderately long snout, small eyes, and a flat disc; tail about as long as body; small inconspicuous, light thorns on upper surface including snout (Ref. 5578).  Medium grey to brownish dorsally, often with dark spots; white or mottled with dusky patches ventrally (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5578&amp;speccode=8938'>Compagno, L.J.V., D.A. Ebert and M.J. Smale 1989 Guide to the sharks and rays of southern Africa. New Holland (Publ.) Ltd., London. 158 p. (Ref. 5578)</reference></dataObject><dataObject>
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	<dc:description>Eastern Atlantic:  off eastern and western South Africa to off Mauritania</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=96993&amp;speccode=8938'>Weigmann, S., M.F.W. Stehmann and R. Thiel 2014 &lt;i&gt;Rajella paucispinosa&lt;/i&gt; n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the western Indian Ocean off South Mozambique, and a revised generic diagnosis. Zootaxa 3847(3):359-387. (Ref. 96993)</reference></dataObject><dataObject>
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	<dc:description>fisheries: commercial</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2683'>Schneider, W. 1990 FAO species identification sheets for fishery purposes. Field guide to the commercial marine resources of the Gulf of Guinea. Prepared and published with the support of the FAO Regional Office for Africa. Rome: FAO. 268 p.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=8938&amp;StockCode=9259</dc:source>
	<dc:description>Oviparous, paired eggs are laid.  Embryos feed solely on yolk (Ref. 50449).  Distinct pairing with embrace.  Young may tend to follow large objects, such as their mother (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=8938'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-8938</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1997-03-16 00:00:00</dcterms:created>
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	<dc:source>leopardus</dc:source>
	<dc:description>Found in deeper shelf and slope waters (Ref. 4426).  Feed on bottom crustaceans, dragonets, rattails, hake, lanternfish, sea pens, cuttlefish and polychaete worms (Ref. 5578).  Feeds on nekton and zoobenthos (Ref. 5578).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7458&amp;speccode=8938'>Ebert, D.A., P.D. Cowley and L.J.V. Compagno 1991 A preliminary investigation of the feeding ecology of skates (Batoidea: Rajidae) off the west coast of southern Africa. S. Afr. J. Mar. Sci. 10:71-81. (Ref. 7458)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>95.0 cm TL (male/unsexed; (Ref. 106604))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=8938'>Weigmann, S. 2016 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
	<dcterms:created>1990-10-18 00:00:00</dcterms:created>
	<dcterms:modified>2024-03-07 00:00:00</dcterms:modified>
	<dc:title></dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2</dc:source>
	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
	<dc:identifier>FB-Habitat-8938</dc:identifier>
	<dataType>http://purl.org/dc/dcmitype/Text</dataType>
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	<agent homepage='' logoURL='' role='recorder'>Crispina B. Binohlan</agent>
	<dcterms:created>1994-03-04 00:00:00</dcterms:created>
	<dcterms:modified>2021-10-15 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=8938</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#LifeCycle</subject>
	<dc:description>bathydemersal; marine; depth range 130 - 1920 m (Ref. 106604)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106604&amp;speccode=8938'>Weigmann, S. 2016 Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. J. Fish Biol. 88(1):1-201. (Ref. 106604)</reference></dataObject>
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