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	<dc:identifier>FB-9128</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9128</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Cichliformes</dwc:Order>
	<dwc:Family>Cichlidae</dwc:Family>
	<dwc:Genus>Tylochromis</dwc:Genus>
	<dwc:ScientificName>Tylochromis leonensis Stiassny, 1989</dwc:ScientificName>
	<commonName xml:lang='fi'>Punaeväseuloja</commonName>
	<synonym relationship='misapplied name'>Pelmatochromis intermedius (non Boulenger, 1916)</synonym>
	<synonym relationship='misapplied name'>Tylochromis intermedius (non Boulenger, 1916)</synonym>
	<dcterms:created>1994-03-07</dcterms:created>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=1880&amp;speccode=9128'>Boulenger, G.A.0 Catalogue of the fresh-water fishes of Africa in the British Museum (Natural History). Volume IV. London. 392 p. (Ref. 1880)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=2990&amp;speccode=9128'>Boulenger, G.A.0 A list of the freshwater fishes of Sierra Leone. Ann. Mag. Nat. Hist. (Ser. 9) 4(19):34-36. (Ref. 2990)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=9128'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5714&amp;speccode=9128'>Stiassny, M.L.J.0 Tylochromis. p. 526-533. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5714)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7378&amp;speccode=9128'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 714-779. In C. Levêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest. Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, 902 p. (Ref. 7378)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12468&amp;speccode=9128'>Baensch, H.A. and R. Riehl0 Aquarien Atlas. Band 4. Mergus Verlag GmbH, Verlag für Natur-und Heimtierkunde, Melle, Germany. 864 p. (Ref. 12468)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=9128'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13331&amp;speccode=9128'>Paugy, D., K. Traoré and P.S. Diouf0 Faune ichtyologique des eaux douces d'Afrique de l'Ouest. p.35-66. In G.G. Teugels, J.F. Guégan and J.J. Albaret (eds.) Biological diversity of African fresh-and brackish water fishes. Geographical overviews presented at the PARADI Symposium, Senegal, 15-20 November 1993. Ann. Mus. R. Afr. Centr., Sci. Zool., 275:177 p. (Ref. 13331)</reference>
	<reference isbn='2-7099-I 340-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=28587&amp;speccode=9128'>Diouf, P.S.0 Les peuplements de poissons des milieux estuariens de l'Afrique de l'Ouest: L'exemple de l'estuaire hyperhalin du Sine-Saloum. Université de Montpellier II. Thèses et Documents Microfiches No.156. ORSTOM, Paris. 267 p. (Ref. 28587)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=9128'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=9128'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=9128'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='3-928819-33-X' url='http://www.fishbase.org/references/FBRefSummary.php?id=52307&amp;speccode=9128'>Lamboj, A.0 The cichlid fishes of Western Africa. Birgit Schmettkamp Verlag, Bornheim, Germany. 255 p. (Ref. 52307)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52345&amp;speccode=9128'>Stiassny, M.L.J.0 &lt;i&gt;Tylochromis&lt;/i&gt;, relationships and the phylogenetic status of the African Cichlidae. Am. Mus. Novit. 2993:1-14. (Ref. 52345)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9128'>Stiassny, M.L.J.0 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52971&amp;speccode=9128'>Regan, C.T.0 A revision of the African cichlid fishes of the genus &lt;i&gt;Tylochromis&lt;/i&gt;. Ann. Mag. Nat. Hist. 5(26):163-169. (Ref. 52971)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=9128'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference>
	<reference isbn='3-928457-13-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=53407&amp;speccode=9128'>Freyhof, J.0 &lt;i&gt;Tylochromis leonensis&lt;/i&gt; Stiassny, 1989. p. 60-61. In A. Konings (ed.) The cichlids yearbook Vol 3. Cichlid press, St. Leon-Rot, Germany, 96p. (Ref. 53407)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57398&amp;speccode=9128'>Teugels, G.G. and D.F.E. Thys van den Audenaerde0 Cichlidae. p. 520-600. In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturelle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57398)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83624&amp;speccode=9128'>Konan, K.F., A. Ouattara, M. Ouattara and G. Gourčne0 Weight-length relationship of 57 fish species of the coastal rivers in South-eastern of Ivory coast. Ribarstvo 65(2):49-60. (Ref. 83624)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=9128'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
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	<dc:title>Biology</dc:title>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<dc:description>Appears to be an exclusively Upper Guinean Province species (Ref. 52346). Inhabits large sandbanks and hardly ever penetrates into small streams (Ref. 53407). Benthic macrophage (Ref. 52346). Feeds primarily upon insects and their larvae, but also accepts plants (Ref. 52307). Oviphilous (Ref. 53407), maternal mouthbrooder that does not pair bond (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=9128'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-9128</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9128</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Diagnosis: 2 scale rows between upper lateral line and anterior part of lower lateral line (Ref. 52346, 53405). High lateral line count (38-42); low gill raker count (13-15 along lower limb of first arch); lower pharyngeal jaw with a short, deep keel that is deeply indented along the ventral margin (Ref. 52346).Description: juveniles gracile, slender-bodied fish that become increasingly deep-bodied and robust as they grow larger; predorsal profile in adults rises more or less smoothly to dorsal fin origin, but a fleshy bulge in interorbital region often interrupts outline (Ref. 52346). Front of head slightly rounded (Ref. 52307). Deepest body depth along back lies at or a little behind origin of dorsal fin; lower jaw more or less along horizontal when mouth is closed, but may become increasingly oblique in larger individuals; lower pharyngeal jaw relatively short and robust; lips well-developed but not noticeably thickened or fleshy (Ref. 52346). Outer and inner jaw teeth unicuspid, pharyngeal jaw teeth molariform (Ref. 53405). First hypobranchial and last ceratobranchial rakers often reduced in size while other ceratobranchial rakers are broad and stout; inferior vertebral apophysis composed of an elongate, caudally straight-edged spine borne the third vertebral centrum (Ref. 52346). Micro-gillrakers present; scales cycloid (Ref. 53405). 2 scale rows between upper lateral line and anterior part of lower lateral line (Ref. 52307, 52346, 53405). 38-42 lateral line scales (Ref. 52346, 53405). Upper branch of lateral line terminates well in advance of end of dorsal fin, usually below 7th-5th ray from free end of fin; terminal canal bearing scales usually does not descend scale rows; dorsal and ventral branches of lateral line on caudal fin extend almost to fin periphery, while median branch is frequently shorter; dorsal fin spines rapidly increasing in length to 5th-6th and then more or less equal in length; anal fin rounded; pectoral fins relatively short and rarely reaching level of spinous anal fin; first branched pelvic ray often produced and filamentous, often reaching beyond spinous anal fin; caudal fin emarginate and in larger individuals finely but uniformily scaled almost to periphery; caudal scaling minimal in juveniles (Ref. 52346).Coloration: No information about color pattern in life (Ref. 52346, 53405). Preserved specimens uniformely light (or sometimes dark) brown (Ref. 52346, 53405), with silvery dots on body scales (Ref. 52307). In specimens smaller than 18-20cm SL upper lip and ethmoidal region greyish (Ref. 52346, 53405), a diffuse opercular blotch present (Ref. 52346), and sides with 6-7 cross bars extending to below midline, which are very distinct in juveniles (juvenile barring), but fewer and less apparent, and not extending below midline in specimens over 18-20cm SL (Ref. 52346, 53405). Dorsal fin with mesh-like patterning of numerous non-occelate pale maculae set upon a smokey grey background (Ref. 52346). Mature males have a prominent interorbital stripe in addition to a strongly marked nape band (Ref. 52346, 53405), and filamentous first branched ray, and much of rest of pelvic fin, dark smokey grey; in male specimens smokey grey anal fin bears faint traces of a few white maculae in proximal field, which is not the case in female specimens; caudal fin bears numerous thin black stripes (Ref. 52346). Anterior parts of anal fin light red with dark spots that are usually visible (Ref. 52307).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9128'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Tobias Musschoot</agent>
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	<dc:description>Dorsal spines (total): 13 - 15; Dorsal soft rays (total): 13 - 16;  Analspines: 3; Analsoft rays: 7; Vertebrae: 28</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52346&amp;speccode=9128'>Stiassny, M.L.J. 1989 A taxonomic revision of the African genus &lt;i&gt;Tylochromis&lt;/i&gt; (Labroidei, Cichlidae); whith notes on the anatomy and relationships of the group. Ann. Mus. R. Afr. Cent. Ser. 8, Sci. Zool., 258:1-161. (Ref. 52346)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Africa:  coastal rivers from the Kogon (Guinea) to the Bia (Sierra Leone) (Ref. 53405).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=5714&amp;speccode=9128'>Stiassny, M.L.J. 1991 Tylochromis. p. 526-533. In J. Daget, J.-P. Gosse, G.G. Teugels and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels; MRAC, Tervuren; and ORSTOM, Paris. Vol. 4. (Ref. 5714)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Grace Tolentino Pablico</agent>
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	<dc:source>leonensis</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>Appears to be an exclusively Upper Guinean Province species (Ref. 52346).  Inhabits large sandbanks and hardly ever penetrates into small streams (Ref. 53407).  Benthic macrophage (Ref. 52346).  Feeds primarily upon insects and their larvae, but also accepts plants (Ref. 52307).</dc:description>
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	<reference isbn='3-928457-13-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=53407&amp;speccode=9128'>Freyhof, J. 1993 &lt;i&gt;Tylochromis leonensis&lt;/i&gt; Stiassny, 1989. p. 60-61. In A. Konings (ed.) The cichlids yearbook Vol 3. Cichlid press, St. Leon-Rot, Germany, 96p. (Ref. 53407)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9128</dc:source>
	<dc:description>21.9 cm SL (male/unsexed; (Ref. 53405))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=53405&amp;speccode=9128'>Teugels, G.G. and D.F.E. Thys van den Audenaerde 2003 Cichlidae. p. 521-600. In D. Paugy, C. Lévêque and G.G Teugels (eds.) The fresh and brackish water fishes of West Africa Volume 2. Coll. faune et flore tropicales 40. Institut de recherche de développement, Paris, France, Muséum national d'histoire naturelle, Paris, France and Musée royal de l'Afrique Central, Tervuren, Belgium, 815p. (Ref. 53405)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Belen Acosta</agent>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-03-07 00:00:00</dcterms:created>
	<dcterms:modified>2024-04-16 00:00:00</dcterms:modified>
	<dc:title>Environment</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9128</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>benthopelagic; freshwater</dc:description>
	<mediaURL></mediaURL>
	<thumbnailURL></thumbnailURL>
	<location></location></dataObject>
</taxon>
</response>