Raja clavata, Thornback ray : fisheries, gamefish

Raja clavata Linnaeus, 1758

Thornback ray
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Native range | All suitable habitat | Point map | Year 2050
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Image of Raja clavata (Thornback ray)
Raja clavata
Immagine di Stergiou, K.I.

Classificazione / Nomi Nomi Comuni | Sinonimi | Catalog of Fishes(Genere, Specie) | ITIS | CoL | WoRMS | Cloffa

Elasmobranchi (squali e razze) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates)
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).
More on author: Linnaeus.

Ambiente: milieu / zona climatica / distribuzione batimetrica / gamma di distribuzione Ecologia

marino demersale; distribuzione batimetrica 5 - 1020 m (Ref. 114953). Subtropical; 70°N - 38°S, 25°W - 50°E

Distribuzione Territories | Aree FAO | Ecosistemi | Presenze | Mappa dei ritrovamenti | Introduzioni | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Lunghezza alla prima maturità / Dimensione / Peso / Età

Maturità: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL maschio/sesso non determinato; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL maschio/sesso non determinato; (Ref. 9987); peso massimo pubblicato: 18.0 kg (Ref. 35388); Età massima riportata: 15 anni (Ref. 41305)

Breve descrizione Chiavi di identificazione | Morfologia | Morfometria

Spine dorsali (totale) : 0; Spine anali: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).
Body shape (shape guide): other; Cross section: flattened.

Biologia     Glossario (es. epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Ciclo vitale e comportamento di accoppiamento Maturità | Riproduzione | Deposizione | Uova | Fecondità | Larve

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Riferimento principale Caricare le referenze | Bibliografia | Coordinatore : McEachran, John | Collaboratori

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

Stato della Lista Rossa IUCN (Ref. 130435: Version 2025-2 (Global))

  Near Threatened (NT) (A2bd); Date assessed: 15 April 2023

CITES

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Minaccia per l'uomo

  Harmless





Usi umani

Pesca: commerciale; Pesce da pesca sportiva: si
FAO - pesca: landings search; Publication: search | FishSource | Sea Around Us

Informazioni ulteriori

Ecologia trofica
Articoli alimentari (prede)
Alimentazione
Consumo di cibo
Razioni alimentari
Predatori
Ecologia
Ecologia
Dinamica della popolazione
Parametri di crescita
Età massima / dimensioni massime
Rapporto lunghezza-peso.
Lunghezza-lunghezza rel.
Lunghezza-frequenze
Conversione di massa
Reclutamento
Abbondanza
Ciclo di vita
Riproduzione
Maturità
Maturità/Gli occhielli sono relativi.
Fecondità
Deposizione
Aggregazioni riproduttive
Uova
Sviluppo delle uova
Larve
Dinamica popolazioni larvali
Distribuzione
Territories
Aree FAO
Ecosistemi
Presenze
Introduzioni
BRUVS - Video
Anatomia
Area branchiale
Cervello
Otolite
Fisiologia
Composizione corporea
Nutrienti
Consumo di ossigeno
Tipo di nuoto
Velocità di nuoto
Pigmenti visivi
Suono del pesce
Malattie e parassiti
Tossicità (LC50)
Genetica
Genome
Genetica
Eterozigosi
Ereditarietà
Diversità genetica
Legato all'uomo
Sistemi di acquacoltura
Profili di acquacoltura
Varietà
Casi di Ciguatera
Francobolli, monete, materiale vario.
Raggiungimento
Collaboratori
Tassonomia
Nomi Comuni
Sinonimi
Morfologia
Morfometria
Immagini
Bibliografia
Bibliografia

Strumenti

Quiz organico | Libro elettronico | Giuda pratica | Chiavi di identificazione | Generatore frequenze di lunghezza | Strumento Parametri Biologici | Mappa dei ritrovamenti | Cattura-MSY |

Rapporti speciali

Controllo della manutenzione dell'acquario | Verificare la presenza di schede informative sulle specie | Verifica delle schede informative sull'acquacoltura

Scarica XML

Pagina riassuntiva | Dati puntuali | Nomi Comuni | foto

Fonti Internet

AFORO (otoliths) | Aquatic Commons | BHL | Cloffa | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes: Genere, Specie | DiscoverLife | DORIS | ECOTOX | FAO - pesca: landings search; Publication: search | Faunafri | Fishipedia | Fishtrace | GloBI | Google Books | Google Scholar | Google | IGFA World Record | Database Nazionali | OneZoom | Open Tree of Life | Otolith Atlas of Taiwan Fishes | Acquari pubblici | PubMed | Reef Life Survey | Socotra Atlas | TreeBase | Tree of Life | Wikipedia: Go, ricerca | World Records Freshwater Fishing | Zoobank | Record zoologico

Stime basate su modelli

Indice di diversità filogenetica
(Fonte Biblio. 82804) 		PD50 = 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight a=0.00204 (0.00179 - 0.00233), b=3.25 (3.22 - 3.28), in cm total length, based on LWR estimates for this species (Ref. 93245).
Livello trofico
(Fonte Biblio. 69278) 		3.8   ±0.2 se; based on diet studies.
Generation time 7.8 (5.4 - 10.4) years. Estimated as median ln(3)/K based on 16 growth studies.
Resilienza
(Fonte Biblio. 120179) 		Basso, tempo minimo di raddoppiamento della popolazione 4.5 - 14 anni (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Rate of population increase Prior r = 0.18, 95% CL = 0.12 - 0.26, Based on 4 full stock assessments.
Vulnerabilità della pesca
(Ref. 59153) 		High vulnerability (60 of 100). 🛈
Vulnerabilità climatica
(Ref. 125649) 		High vulnerability (60 of 100). 🛈
Categoria di prezzo
(Ref. 80766) 		Medium.
Nutrienti
(Ref. 124155) 		Calcium = 9.22 [1.18, 176.74] mg/100g; Iron = 0.628 [0.060, 7.651] mg/100g; Protein = 17.2 [14.4, 19.5] %; Omega3 = 0.571 [0.265, 1.295] g/100g; Selenium = 39.7 [7.5, 183.1] μg/100g; VitaminA = 5.22 [0.40, 59.91] μg/100g; Zinc = 0.47 [0.03, 5.30] mg/100g (wet weight); based on nutrient studies.