Raja clavata, Thornback ray : fisheries, gamefish

Raja clavata Linnaeus, 1758

Thornback ray
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Image of Raja clavata (Thornback ray)
Raja clavata
Bild av Stergiou, K.I.

Klassificering / Namn Populärnamn | synonymer | Catalog of Fishes(Släkte, Arter) | ITIS | CoL | WoRMS | Cloffa

Elasmobranchii (hajar och rockor) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates)
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).
More on author: Linnaeus.

Miljö: miljö / Klimatzon / djupintervall / distributionsområde Ekologi

marina bottenlevande; djupintervall 5 - 1020 m (Ref. 114953). Subtropical; 70°N - 38°S, 25°W - 50°E

Utbredning Territorier | FAO områden | Ekosystem | Förekomster | Prickkarta | Utplanteringar | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Längd vid första mognad / Storlek / Vikt / Ålder

Könsmognad: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL hane/ej könsbestämd; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL hane/ej könsbestämd; (Ref. 9987); publicerad maxvikt: 18.0 kg (Ref. 35388); rapporterad maxålder: 15 år (Ref. 41305)

Kort beskrivning Bestämningsnycklar | Morfologi | Morfometri

Taggstrålar i ryggfenan (totalt) : 0; Taggstrålar i analfenan: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).
Body shape (shape guide): other; Cross section: flattened.

Biologi     Ordlista (t.ex. epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Livscykel och parningsbeteende Könsmognad | Reproduktion | Lek | Ägg | Fekunditet | Larver

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Huvudreferens Ladda upp dina referenser | referenser | Koordinator : McEachran, John | Medarbetare

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

Status på IUCN:s rödlista (Ref. 130435: Version 2025-2 (Global))

  Near Threatened (NT) (A2bd); Date assessed: 15 April 2023

CITES

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Hot mot människor

  Harmless





Mänskliga användningsområden

Fiskeri: kommersiell; sportfisk: ja
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Ytterligare information

Livscykel
Reproduktion
Könsmognad
Mognad/Gills rel.
Fekunditet
Lek
Lekande aggregat
Ägg
Utveckling av ägg
Larver
Larvdynamik
Utbredning
Territorier
FAO områden
Ekosystem
Förekomster
Utplanteringar
BRUVS - Videor
Anatomi
Gälyta
Hjärna
Otolit
Fysiologi
Kroppssammansättning
Näringsämnen
Syreförbrukning
Typ av simning
Simhastighet
Visuella pigment
Ljud från fisk
Sjukdomar & Parasiter
Toxicitet (LC50)
Genetik
Genom
Genetik
Heterozygositet
Ärftlighet
Genetisk mångfald
Människorelaterad
Vattenbrukssystem
Vattenbruksprofiler
Avelslinjer
Ciguatera-fall
Frimärken, mynt, diverse.
Uppsökande verksamhet
Medarbetare
referenser
referenser

Verktyg

Särskilda rapporter

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Internet-källor

AFORO (otoliths) | Aquatic Commons | BHL | Cloffa | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes: Släkte, Arter | DiscoverLife | DORIS | ECOTOX | FAO - fiskeri: landings search; Publication: search | Faunafri | Fishipedia | Fishtrace | GloBI | Google Books | Google Scholar | Google | IGFA World Record | Nationella databaser | OneZoom | Open Tree of Life | Otolith Atlas of Taiwan Fishes | Offentliga akvarier | PubMed | Reef Life Survey | Socotra Atlas | TreeBase | Tree of Life | Wikipedia: Go, sök | World Records Freshwater Fishing | Zoobank | Zoologiskt register

Uppskattningar baserade på modeller

Index för fylogenetisk mångfald
(Ref. 82804)
PD50 = 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight a=0.00204 (0.00179 - 0.00233), b=3.25 (3.22 - 3.28), in cm total length, based on LWR estimates for this species (Ref. 93245).
Trofisk nivå
(Ref. 69278)
3.8   ±0.2 se; based on diet studies.
Generation time 7.8 (5.4 - 10.4) years. Estimated as median ln(3)/K based on 16 growth studies.
Resiliens
(Ref. 120179)
Låg, lägsta populationsfördubblingstid 4,5-14 år (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Rate of population increase Prior r = 0.18, 95% CL = 0.12 - 0.26, Based on 4 full stock assessments.
Sårbarhet i fisket
(Ref. 59153)
High vulnerability (60 of 100). 🛈
Sårbarhet för klimatet
(Ref. 125649)
High vulnerability (60 of 100). 🛈
Priskategori
(Ref. 80766)
Medium.
Näringsämnen
(Ref. 124155)
Calcium = 9.22 [1.18, 176.74] mg/100g; Iron = 0.628 [0.060, 7.651] mg/100g; Protein = 17.2 [14.4, 19.5] %; Omega3 = 0.571 [0.265, 1.295] g/100g; Selenium = 39.7 [7.5, 183.1] μg/100g; VitaminA = 5.22 [0.40, 59.91] μg/100g; Zinc = 0.47 [0.03, 5.30] mg/100g (wet weight); based on nutrient studies.